A Review of the Genus Eunice - Smithsonian Institution Libraries

168 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
1826); he cited E. harassii as a synonym. Risso's species is not
identifiable (see below); Heider's description fits E. harassii,
thus his detailed anatomical work is assumed to be of this
species.
Eunice harassii is a member of B-2, but differs from most
species in that group by having brown aciculae and subacicular
hooks rather than black ones as usual in that group. It is listed
with similar species in Tables 27 and 31. One unique character
state of this species appear to be the broadly attached, flattened,
nearly leaf-shaped ventral cirri in posterior setigers. The
detailed branchial distribution may also be characteristic, in
that the number of filaments is reduced from middle segments,
but branchiae are retained through the rest of the body.
84. Eunice havaica Kinberg, 1865
FIGURE 54e-g; TABLES 46-48
Eunice havaica Kinberg, 1865: 562; 1910:41, pi. 15: fig. 14.
Eunice antennata.—Hartman, 1948:78 [in part, not Eunice antennata Lamarck,
1818].
MATERIAL EXAMINED.—14 syntypes, RM 430, Honolulu,
Hawaii, Eugenie Expedition 1087-89.
COMMENTS ON MATERIAL EXAMINED.—All but one of the
15 specimens of the original material of E. havaica belong to
this species; the last one differs from the others at the specific
level and is described above as E. eugeniae. All the syntypes
are poorly preserved and most are anterior fragments only. Due
to the poor preservation parapodial shape can be described only
tentatively.
DESCRIPTION.—Complete syntype of unknown sex, with 78
setigers; total length 25 mm; maximal width 1 mm at setiger
10; length through setiger 10,4 mm.
Prostomium (Figure 54h) distinctly shorter than peristomium,
about as wide as peristomium, less than l /2 as deep as
peristomium. Prostomial lobes frontally obliquely truncate,
dorsally flattened; median sulcus shallow. Eyes barely visible
well behind bases of A-I. Antennae in horseshoe, evenly
spaced, similar in thickness. Ceratophores ring-shaped in all
antennae, without articulations. Ceratostyles tapering, with up
to 22 moniliform articulations in A-III, articulated to bases. A-I
to middle of first peristomial ring; A-I I to setiger 2; A-I 11 to
setiger 3. Peristomium cylindrical. Separation between rings
distinct on all sides; anterior ring 3 A of total peristomial length.
Peristomial cirri to frontal margin of peristomium, slender, with
9-10 moniliform articulations.
Summary maxillary formula for 2 syntypes 1+1, 7+7, 7+0,
6-8+8, and 1+1. Max III long, located behind left Mx II. Mx VI
missing.
Branchiae present, pectinate, about as long as notopodial
cirri, reduced in mid-body region, erecL Branchiae from setiger
6 to setiger 78. Branchiae present to near posterior end, on more
than 65% of total number of setigers. In complete syntype
branchial occurrence interrupted between setigers 28 and 48
leaving median segments abranchiate. Maximum number of
filaments 5 at about setiger 20; maximum number of filaments
in posterior branchiated region 3. Stems slender, tapering,
shorter than notopodial cirri. Filaments slender, digitiform,
shorter than notopodial cirri.
Neuropodial acicular lobes distally apparently truncate;
aciculae emerging at midline. Prc- and postsetal lobes low,
transverse folds. Anterior and posterior ventral cirri digitiform,
basally inflated between setigers 5 and 40. Inflated bases ovate;
narrow tips tapering. Notopodial cirri tapering, with up to 6
articulations at least in first 25 setigers; articulations indistinct
posteriorly; far posterior notopodial cirri without articulations.
Limbate setae slender. Pectinate setae flat, tapering. Both
marginal teeth slightly longer than other teeth, with -15 teeth.
Shafts of compound falcigers (Figure 540 distally inflated,
usually marginally smooth. Appendages short with large head,
bidentatc or tridcntatc (Figure 54c). Proximal and distal teeth of
same size. Proximal teeth narrowly triangular, directed laterally.
Distal teeth distinctly bent, split in some hooks, forming
distinctly tridcntatc appendages. Guards symmetrically bluntly
pointed; mucros absent. Pscudocompound falcigers and compound
spinigers absent. Aciculae yellow, tapering to bluntly
conical tips, straight; cross-sections round. Separation between
core and sheath indistinct in both aciculae and subacicular
hooks. Subacicular hooks (Figure 54g) yellow, tridentate with
teeth in a crest. Hooks first present from setiger 19-23, present
in all setigers thereafter, always single (except for replacements).
Main fang large, curved. Smaller fangs decreasing
evenly in size.
UNKNOWN MORPHOLOGICAL FEATURES.—Pygidium
anal cirri.
EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—
None.
CHARACTERS USED IN PREPARATION OF KEY NOT
SCORED.—Inappropriate Characters: 56, 58, 59. Unknown
Characters: 39,40.
ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—
None.
REMARKS.—Eunice havaica was considered a synonym of
E. antennata by Hartman (1948). It is listed in Tables 46 and
48. Of the species listed in Table 48 only E. havaica and E.
stigmatura have complete cinctures marking off the separation
between the two peristomial rings. The compound falcigers
have both bi- and tridentate appendages in E. havaica and only
bidentate appendages in E. stigmatura.
Hartman (1948:78) indicated that she had seen only one of
the syntypes of this species, but later indicated a range of values
for first occurrence of subacicular hooks, showing that she must
have examined at least several specimens. The specimens are
very soft, making study of variability difficult.
and