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A Review of the Genus Eunice - Smithsonian Institution Libraries

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166 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY<br />

remaining inflated through rest <strong>of</strong> body. Basal inflation very<br />

large, nearly spherical, very small tapering tips present in all<br />

setigers. Notopodial cirri basally inflated, distally tapering,<br />

without articulations.<br />

Limbate setae tapering, gently geniculate, frayed. Pectinate<br />

setae (Figure 53i) numbering 5 or more in a fascicle. Shafts<br />

wide, flat Blades tapering, flat. One marginal tooth clearly<br />

longer than o<strong>the</strong>r teeth, with -14 teeth. Shafts <strong>of</strong> compound<br />

falcigers (Figure 53h) distally inflated, marginally smooth.<br />

Appendages very slender, tapering, with large heads, bidentate.<br />

Proximal teeth about as large as distal teeth, sharply directed<br />

laterally. Distal teeth flattened, nearly geniculate and directed<br />

laterally. Guards asymmetrically bluntly pointed, marginally<br />

smooth; mucros absent. Pseudocompound falcigers and compound<br />

spinigers absent Aciculae numbering 1 to 3, dark,<br />

tapering, straight; cross-sections round. Notopodial aciculae<br />

paired, dark. Subacicular hooks (Figure 53j) dark, bidentate.<br />

Hooks first present from setiger 32, present in all setigers<br />

<strong>the</strong>reafter, paired in some setigers. Hooks smoothly tapering to<br />

small head. Proximal teeth larger than distal teeth; both teeth<br />

directed obliquely distally. Guards short and blunt.<br />

UNKNOWN MORPHOLOGICAL FEATURES.—Jaw structure;<br />

pygidium and anal cirri.<br />

EXPECTED STATES OF SELECTED UNKNOWN FEATURES —<br />

Mx III short and forming part <strong>of</strong> distal arc with left Mx IV.<br />

CHARACTERS USED IN PREPARATION OF KEY NOT<br />

SCORED.—Inappropriate Characters: 22, 56, 60. Unknown<br />

Characters: 13, 14, 39,40,42, 74, 78.<br />

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—<br />

None.<br />

REMARKS.—The specimen may have been highly colored; a<br />

ghost <strong>of</strong> dark pigmentation remains. The specimen may have<br />

been dark brown or red, with white circular spots scattered<br />

densely over <strong>the</strong> anterior end. A white band may have been<br />

present.<br />

As far as known, this species has never been reported since<br />

it was briefly described by Baird (1869); it belongs to group<br />

B-2 and is listed with similar species in Tables 27 and 28. It is<br />

<strong>the</strong> only species, o<strong>the</strong>r than E. aphroditois, listed in Table 28<br />

that has <strong>the</strong> ventral cirri basally inflated through <strong>the</strong> posterior<br />

end. It differs from that species mainly in that it has far fewer<br />

branchial filaments and in <strong>the</strong> proportions <strong>of</strong> <strong>the</strong> prostomium.<br />

Nothing is known about its habitat o<strong>the</strong>r than that it was<br />

collected somewhere in <strong>the</strong> Indian Ocean/Indonesian region,<br />

presumably in shallow water.<br />

83. <strong>Eunice</strong> harassii Audouin and Milne Edwards, 1833<br />

FIGURE 54a-d; TABLES 27,31<br />

<strong>Eunice</strong> harassii Audouin and Milne Edwards, 1833:215-218, pi. 11: figs.5-7,<br />

10, 11; 1834:141-144, pi. 3: figs. 5-7. 10, 11.—Quatrefages, 1866:307-<br />

309, pi. 10: fig. 3.<br />

Eunke pimctata.—Heider, 1925:55-89, figs. 1, 2, 7-16 [not Leodice<br />

punctata Risso, 1826].<br />

MATERIAL EXAMINED.—One specimen, MNHN, Paris<br />

A.l(R.)-1868-no. 46a. lies Chausey, Quatrefages.<br />

COMMENTS ON MATERIAL EXAMINED.—The type has been<br />

lost; justification for using <strong>the</strong> current specimen to represent <strong>the</strong><br />

species is given below. The jaws are half everted and <strong>the</strong><br />

maxillary formula given is incomplete because detailing <strong>the</strong><br />

formula would destroy <strong>the</strong> jaws completely.<br />

DESCRIPTION.—Specimen incomplete, <strong>of</strong> unknown sex,<br />

with 63 setigers; length 22 mm; maximal width 2.5 mm at<br />

setiger 10; length through setiger 10, 5 mm. Body cylindrical.<br />

Prostomium less than x li as thick as peristomium, narrow.<br />

Prostomium (Figure 54a) distinctly shorter and narrower than<br />

peristomium, less than x li as deep as peristomium. Prostomial<br />

lobes frontally obliquely truncate, dorsally flattened; median<br />

sulcus deep. Eyes at bases <strong>of</strong> A-I, faded. Antennae in shallow<br />

horseshoe, evenly spaced, similar in thickness. Ccratophorcs<br />

ring-shaped in all antennae, without articulations. Ceratostyles<br />

tapering, with up to 5 indistinct, cylindrical articulations in<br />

A-I 11. A-I to posterior pcristomial ring; A-I I to setiger 2; A-I 11<br />

to setiger 4. Peristomium cylindrical. Separation between rings<br />

distinct on all sides; anterior ring 4 /s <strong>of</strong> total pcristomial length.<br />

Pcristomial cirri to middle <strong>of</strong> anterior pcristomial ring, slender<br />

and tapering, without articulations.<br />

Incomplete maxillary formula 1 + 1, +, 6+0, 3+8, 1+1. Mx<br />

VI missing. Mx III long, but part <strong>of</strong> distal arc with left Mx IV.<br />

Branchiae (Figure 54d) present, pectinate, distinctly longer<br />

than notopodial cirri, not reduced in mid-body region, erect.<br />

Branchiae from setiger 4 to end <strong>of</strong> fragment. First 3 branchiae<br />

single filaments; all o<strong>the</strong>rs pectinate with maximally 10<br />

filaments at about setiger 15; last branchia present, on setiger<br />

63, with 2 filaments only. Stems tapering, shorter than<br />

notopodial cirri. Filaments long, digitiform, as long as<br />

notopodial cirri where best developed.<br />

Anterior neuropodial acicular lobes truncate; posterior<br />

acicular lobes triangular; aciculae emerging at midline. Pre-and<br />

postsetal lobes low, transverse folds. First 5 ventral cirri thick,<br />

tapering. Ventral cirri basally inflated from setiger 6 through<br />

about setiger 35; inflated base ovate; narrow lips tapering.<br />

Posterior ventral cirri broadly transversely attached; enfolding<br />

lower edge <strong>of</strong> parapodia, nearly scoop-shaped. Anterior<br />

notopodial cirri basally slightly inflated, retaining same size in<br />

all setigers, but less distinctly inflated in posterior setigers,<br />

without articulations.<br />

Limbate setae slender, marginally smooth. Pectinate setae<br />

tapering, flat. One marginal tooth longer than o<strong>the</strong>r teeth, with<br />

-12 teeth. Shafts <strong>of</strong> compound falcigers (Figure 54c) distally<br />

tapering, marginally smooth. Appendages relatively short,<br />

tapering with large, distinct heads, bidentate. Proximal teeth as<br />

large as distal teeth, triangular, directed laterally. Distal teeth<br />

distinctly bent, sharply tapering. Guards asymmetrically<br />

bluntly pointed, marginally smooth; mucros absent. Pseudocompound<br />

falcigers and compound spinigers absent. Aciculae<br />

paired; anterior aciculae honey colored, becoming darker<br />

brown in posterior setigers, distally tapering to sharp points,

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