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A manual of rice seed health testing - IRRI books - International Rice ...

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Red flour beetle or bran bug<br />

Scientific name: Tribolium castaneum<br />

(Herbst)<br />

Common name: red flour beetle or bran<br />

bug<br />

Quarantine status: primary pest<br />

Stage <strong>of</strong> entrance or attachment to<br />

<strong>seed</strong>: adult, larva, and egg<br />

The body <strong>of</strong> the adult is 3-5 mm long<br />

and shiny red brown to blackish<br />

brown. It may be confused with the<br />

confused flour beetle. The red flour<br />

beetle’s diagnostic features are a<br />

relatively flat body with a broaderthan-long<br />

pronotum, and parallelsided<br />

abdomen; 11-segmented antennae<br />

with a 3-segmented pronounced<br />

club; a head lacking a ridge<br />

above the eyes; and eyes separated<br />

ventrally by a space less than two<br />

times the diameter <strong>of</strong> the eye. The<br />

adult lives 335-540 d.<br />

Each female lays 300-500 eggs on<br />

the outside <strong>of</strong> grains, attaching the<br />

eggs with sticky glue.<br />

Larvae feed both outside and inside<br />

the grain. The larva has a<br />

prominent head, three pairs <strong>of</strong> distinct<br />

legs, forked abdomen tip, and<br />

an elongated and cylindrical white<br />

body tinged with yellow. The life<br />

cycle <strong>of</strong> the red flour beetle lasts<br />

4-6 wk but may decrease to 20 d under<br />

optimum conditions.<br />

The length <strong>of</strong> the life cycle depends<br />

on the availability <strong>of</strong> food.<br />

The pest is omnivorous and cannibalistic.<br />

It is more cold-susceptible<br />

than the cold-hardy Cryptolestes. The<br />

adult is a strong flier.<br />

Confused flour beetle<br />

Scientific name: Tribolium confusurn<br />

(Jacquelin du Val)<br />

Common name: confused flour beetle or<br />

bran bug<br />

Quarantine status: primary pest<br />

Stage <strong>of</strong> entrance or attachment to<br />

<strong>seed</strong>: adult, larva, and egg<br />

The adult beetle is 3-4 mm long and<br />

shiny reddish brown. It may be confused<br />

with the red flour beetle. Diagnostic<br />

features include a flat, ovalshaped<br />

body; 11-segmented antennae<br />

gradually thickening toward the<br />

apex, forming a poorly differentiated<br />

5-segmented club; a head with a<br />

ridge above the eyes; and eyes<br />

separated ventrally by a space three<br />

times the diameter <strong>of</strong> the eye. The<br />

adult lives 1 yr.<br />

Females lay 450-500 eggs on grain.<br />

Larvae feed externally on the grains.<br />

Larvae have prominent heads, the<br />

pairs <strong>of</strong> distinct legs, forked abdomen<br />

tips, and are pale yellow. The<br />

life cycle is 4-6 wk.<br />

The confused flour beetle prefers<br />

to feed on the grain embryo but can<br />

complete development exclusively<br />

on <strong>seed</strong>borne fungi. It is more coldhardy<br />

than the red flour beetle and is<br />

a weak flier. Both adults and larvae<br />

are omnivorous and cannibalistic.<br />

Insect pests 73

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