A manual of rice seed health testing - IRRI books - International Rice ...
A manual of rice seed health testing - IRRI books - International Rice ...
A manual of rice seed health testing - IRRI books - International Rice ...
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CHAPTER 11<br />
Seed treatment<br />
J.K. Misra. T.W. Mew, and C.C. Huelma<br />
Seed treatment is physical or chemical<br />
<strong>seed</strong> therapy. Seed treatment<br />
dates back at least to the Roman empire,<br />
when wine and crushed cypress<br />
leaves were used to treat <strong>seed</strong>s<br />
for better harvest.<br />
Today, <strong>seed</strong> treatment is important<br />
in pest management. In western<br />
countries and Japan, where mechanized<br />
farming is practiced, <strong>seed</strong><br />
treatment has become part <strong>of</strong> normal<br />
farm procedures. Seed treatment<br />
revolutionized production<br />
(Nakamura 1986, Bowling 1986) by<br />
controlling many pests and pathogens<br />
and by repelling birds. As new<br />
technologies and chemicals become<br />
available, the dimensions <strong>of</strong> <strong>seed</strong><br />
treatment change significantly.<br />
Seeds are treated<br />
in the early stages <strong>of</strong> their germination<br />
and establishment to protect<br />
them from pests and pathogens,<br />
to promote better <strong>seed</strong>ling stands,<br />
to minimize yield loss,<br />
to maintain and improve <strong>seed</strong><br />
quality, and<br />
to avoid introduction and spread<br />
<strong>of</strong> harmful organisms.<br />
Advantages <strong>of</strong> <strong>seed</strong><br />
treatment<br />
Among the many advantages <strong>of</strong> <strong>seed</strong><br />
treatment are that it<br />
protects <strong>seed</strong>s and <strong>seed</strong>lings from<br />
pests and pathogens in the early<br />
stages <strong>of</strong> germination and establishment,<br />
is easier than plant treatment and<br />
can be done indoors with or without<br />
machinery,<br />
is unaffected by weather conditions,<br />
requires less chemicals than does<br />
plant treatment,<br />
pollutes the environment less than<br />
does plant treatment,<br />
does not result in much development<br />
<strong>of</strong> resistance in insect pests (although<br />
fungi are developing resistance<br />
to fungicides), and<br />
uses biocides that affect only targeted<br />
organisms and not the other<br />
beneficial entities in the soil.<br />
Limitations <strong>of</strong> <strong>seed</strong><br />
treatment<br />
Advantages <strong>of</strong> <strong>seed</strong> treatment outweigh<br />
the limitations which include<br />
lack <strong>of</strong> effective broad-spectrum<br />
systemic fungicides for internally<br />
<strong>seed</strong>borne fungi and for fungal resting<br />
structures;<br />
no protection beyond the early<br />
growth or <strong>seed</strong>ling stages;<br />
no potent biocides that protect the<br />
<strong>seed</strong>s from rodents, nematodes, insects,<br />
slugs, and birds; and<br />
chemicals do not continue to adhere<br />
to the <strong>seed</strong> surface, do not<br />
maintain the desired level during<br />
germination, and do not protect germinating<br />
<strong>seed</strong>s and <strong>seed</strong>lings from<br />
<strong>seed</strong>borne and soilborne pests and<br />
pathogens.<br />
Methods<br />
Seeds are treated by chemical, physical,<br />
or both methods. Aspects <strong>of</strong> general<br />
<strong>seed</strong> treatment are presented by<br />
Jeffs (1986), Neergaard (1979), and<br />
Anselme (1988). Information about<br />
<strong>rice</strong> <strong>seed</strong> treatment for quarantine<br />
purposes is presented here.<br />
Physical <strong>seed</strong> treatment is any<br />
method that does not employ chemicals.<br />
Normally, heat treatment is accomplished<br />
using hot water, dry<br />
heat, or steam.<br />
Heat treatment kills <strong>seed</strong>borne<br />
pests and pathogens without injury<br />
to most germplasm. Heat therapy<br />
may be injurious to and should be<br />
used cautiously with old or damaged<br />
<strong>seed</strong>s and varieties which have<br />
low heat tolerance. Japonica varieties<br />
are generally more heat-sensitive<br />
than indica varieties.<br />
Hot water treatment at 52-57 °C<br />
for 15 min eradicates <strong>seed</strong>borne<br />
Aphelenchoides besseyi.<br />
Heat therapy is <strong>of</strong>ten given in<br />
conjunction with chemical treatment.<br />
Biocidal chemicals (fungicides,<br />
bactericides, insecticides,<br />
nematicides) are applied as liquids<br />
or powders. Depending on the type<br />
<strong>of</strong> <strong>seed</strong> and the nature <strong>of</strong> the chemical,<br />
biocides may be applied as dust<br />
treatments, fumigants, wet treatments<br />
(steep, sprinkle, quick wet,<br />
and slurry), oil fungicide treatments,<br />
and by pelleting. Since no broadspectrum<br />
fungicide exists, mixtures<br />
<strong>of</strong> fungicides are commonly used.<br />
In Japan, the following chemicals<br />
are used to control blast, bakanae,<br />
and brown spot <strong>rice</strong> diseases<br />
(Nakamura 1986):<br />
thiram 20% and benomyl 20%<br />
(benlate T);<br />
thiram 30% and thiophanatemethyl<br />
50% (Homai);<br />
thiram 10% and thiophanatemethyl<br />
10% (Homai coat);<br />
captan 30% and kasugamycin 3%<br />
(kasumin C); and<br />
copper sulfate, basic 29%, and<br />
oxine-copper 20% (oxybordeau).<br />
Chemicals used for <strong>seed</strong> treatment<br />
in the USA are<br />
captan,<br />
thiram,<br />
maneb,<br />
chloroneb,<br />
difolatan ( cis --N-((l,l,2,2-<br />
tetrachloroethyl) thio)4-<br />
cyclohexene-1,2, dicarboximide),<br />
Vitavax (2,3-dihydro-6-methyl-5-<br />
phenyl carbamoyl-1,4-oxathiin),