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SOUVENIR COMMITTEE - Birbal Sahni Institute of Palaeobotany

SOUVENIR COMMITTEE - Birbal Sahni Institute of Palaeobotany

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Training Programme on “PALYNOLOGY IN FOSSIL FUEL EXPLORATION”<br />

[ September 10-17, 2012 ]<br />

INTRODUCTION TO DINOFLAGELLATES: CYST MORPHOLOGY AND TERMINOLOGY<br />

N.C. MEHROTRA<br />

<strong>Birbal</strong> <strong>Sahni</strong> <strong>Institute</strong> <strong>of</strong> <strong>Palaeobotany</strong>, 53, University Road, Lucknow<br />

E-mail: mehrotrabsip@rediffmail.com, director@bsip.res.in<br />

The din<strong>of</strong>lagellates are in today's environments an extremely diverse group <strong>of</strong> protists. The only<br />

din<strong>of</strong>lagellates <strong>of</strong> importance to paleopalynogists are those which have a complex life cycle with a thin<br />

walled, motile, characteristically flagellate stage usually called the theca or thecate stage, and a<br />

nonmotile, thick-walled encysted stage, the cyst. Sporopolleninous cysts <strong>of</strong> din<strong>of</strong>lagellates are common<br />

from Triassic rocks to present, mostly in marine environments, but also in sediments deposited in fresh<br />

and brackish water. Thecae <strong>of</strong> din<strong>of</strong>lagellates <strong>of</strong> the order Peridiniales are divided into several series <strong>of</strong><br />

more or less polygonal units, the plates. The lines <strong>of</strong> separation between plates and sutures, and the sum<br />

<strong>of</strong> the characteristics <strong>of</strong> the plates (i.e., their number, size, arrangement into series, shape, mutual<br />

contacts) constitute the tabulation. Certain aspects <strong>of</strong> the tabulation (number <strong>of</strong> plates in each series) may<br />

be abbreviated in a tabulation formula (or plate formula) using the form and symbols suggested by<br />

K<strong>of</strong>oid. An unit area on the cyst corresponding to a thecal plate is a paraplate. Paraplates are delineated<br />

by parasutures (i.e., linear features which correspond in position to thecal sutures), or by nonparasutural<br />

features (e.g., intratabular processes or process groups) that are positioned in areas corresponding to the<br />

more central parts <strong>of</strong> the thecal plates. The sum <strong>of</strong> the characteristics <strong>of</strong> the paraplates constitutes the<br />

paratabulation formula. A paratabulation formula gives the number <strong>of</strong> evident paraplates in each series,<br />

but it does not include information on the morphological expression or on the size and mutual<br />

relationships <strong>of</strong> the evident paraplates.<br />

<strong>Birbal</strong> <strong>Sahni</strong> <strong>Institute</strong> <strong>of</strong> <strong>Palaeobotany</strong>, Lucknow<br />

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