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Landscapes Forest and Global Change - ESA - Escola Superior ...

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J.L. Hernández-Stefanoni et al. 2010. Effects of l<strong>and</strong>scape structure <strong>and</strong> st<strong>and</strong> age on species richness <strong>and</strong> biomass<br />

643<br />

in DBH (diameter at breast (1.3 m) height), hereafter referred to as adults, were sampled in a<br />

200 m 2 plot; whereas 1-5 cm DBH woody plants, hereafter referred to as juveniles, were<br />

sampled in a nested 50 m 2 subplot. In each site, we identified all woody plants, <strong>and</strong> measured<br />

their diameter <strong>and</strong> height. The number of adults, juveniles <strong>and</strong> all woody plant species was<br />

computed. To calculate above-ground biomass, two equations were employed: one for<br />

individuals ≥ 10 cm in DBH (Cairns et al, 2003) <strong>and</strong> the other for individuals < 10 cm in DBH<br />

(Hughes et al, 2000).<br />

2.5 Data analysis<br />

Multiple regression <strong>and</strong> variation partitioning methods (Bocard et al, 2004) were used to<br />

quantify the effects of l<strong>and</strong>scape structure <strong>and</strong> spatial dependence on species density <strong>and</strong><br />

biomass. First, a model using l<strong>and</strong>scape structure <strong>and</strong> st<strong>and</strong> age variables was fit to the response<br />

variables using multiple regression. Then, a multiple regression model using spatial variables<br />

derived from a principal coordinate of neighbor matrices (PCNM) analysis of plot locations was<br />

fit to the response variables. Finally, the two models were combined into an overall regression<br />

model <strong>and</strong> variation partitioning was performed to determine the relatively importance of<br />

l<strong>and</strong>scape structure variables, st<strong>and</strong> age, pure spatial dependence <strong>and</strong> shared variation on species<br />

density <strong>and</strong> biomass.<br />

PCNM analysis <strong>and</strong> multiple regressions were used to identify l<strong>and</strong>scape structure <strong>and</strong><br />

st<strong>and</strong> age variables related to response variables at different spatial scales (Bocard et al, 2004).<br />

First, we partitioned the spatial model for each response variable into several additive<br />

submodels, <strong>and</strong> run a variogram analysis of significant PCNM vectors to identify their scale <strong>and</strong><br />

assign them to one of three groups: very board scale (distances of 8001 to 10500 m), broad scale<br />

(distances of 2001 to 8000 m), <strong>and</strong> local scale (distances of 0 to 2000 m). Then, we calculated<br />

predicted values of species density <strong>and</strong> biomass corresponding to each spatial submodel. Finally,<br />

a multiple regression model using st<strong>and</strong> age <strong>and</strong> l<strong>and</strong>scape structure metrics was fit to the<br />

predicted response variables for each submodel (very broad, broad <strong>and</strong> local scale).<br />

3. Results<br />

The l<strong>and</strong> cover thematic map of the study area is shown in Figure 1. This l<strong>and</strong>scape<br />

covers a total area of 37 242 ha; 94.3% is covered by forest in any of the four vegetation classes,<br />

<strong>and</strong> only 5.7% is covered by agriculture, urban areas <strong>and</strong> roads. The overall accuracy calculated<br />

for the map was 75.6%, <strong>and</strong> the Kappa index was 0.7.<br />

Figure 1: Location <strong>and</strong> l<strong>and</strong> cover map of the study area obtained from a supervised classification.<br />

<strong>Forest</strong> <strong>L<strong>and</strong>scapes</strong> <strong>and</strong> <strong>Global</strong> <strong>Change</strong>-New Frontiers in Management, Conservation <strong>and</strong> Restoration. Proceedings of the IUFRO L<strong>and</strong>scape Ecology<br />

Working Group International Conference, September 21-27, 2010, Bragança, Portugal. J.C. Azevedo, M. Feliciano, J. Castro & M.A. Pinto (eds.)<br />

2010, Instituto Politécnico de Bragança, Bragança, Portugal.

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