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Landscapes Forest and Global Change - ESA - Escola Superior ...

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J.O. López-Martínez et al. 2010. Ecological factors influencing beta diversity at two spatial scales in a tropical dry forest<br />

247<br />

Variation in community composition among sites was partitioned using the method of<br />

Borcard et al (1992). Four sets of explanatory variables were considered at the local level:<br />

environmental variables (soil attributes, % stoniness), l<strong>and</strong>scape structure (TECI, SIEI, LPI),<br />

spatial dependence (PCNMs), <strong>and</strong> st<strong>and</strong> age; whereas only two sets of variables were used at<br />

the l<strong>and</strong>scape level: l<strong>and</strong>scape structure <strong>and</strong> spatial dependence. To determine the amount of<br />

variation explained independently by each set of predictor variables, we performed partial<br />

constrained ordination separately for each set of variables at both levels. Due to the length of<br />

gradients, CCA was used at the local level (3.99 SD), while RDA was used at the l<strong>and</strong>scape<br />

scale (2 SD).<br />

3. Results<br />

The l<strong>and</strong> cover thematic map of the study area is shown in Figure 1. The total area<br />

occupied by this l<strong>and</strong>scape was 37,243 ha, 94.8% corresponded to tropical sub-deciduous forest<br />

in any of the four vegetation classes, whereas 5.2% corresponded to agriculture <strong>and</strong> urban areas.<br />

We obtained an overall accuracy in the supervised classification of 73.3% <strong>and</strong> the Kappa index<br />

was of 0.7.<br />

Figure 1: Location <strong>and</strong> l<strong>and</strong> cover map of the study area obtained from a supervised classification.<br />

A total of 22,262 woody individuals belonging to 204 species <strong>and</strong> 52 families were<br />

recorded in 276 sites in 23 l<strong>and</strong>scapes. Of total of sites sampled, 51 belonged at class 1, 77 to<br />

class 2, 83 to class 3 <strong>and</strong> 62 to class 4. The most abundance species was Neomillspaughia<br />

emarginata with 3,421 individuals, whereas 29 species were represented only by a single<br />

individual.<br />

3.1 Local scale.<br />

Detrended correspondence analysis at the local scale showed a large length gradient of species<br />

composition (3.99 sd; Fig. 2a). Vegetation class 4 differed significantly from the other classes.<br />

Canonical correlation analysis showed that total variation explained was 13.79%. Variation<br />

partitioning showed that space is the most important variable explaining 5.95% of species<br />

composition, followed of the soil variables with 5.83% (Table 1). Vegetation class 1 was<br />

negatively associated with organic matter (SOM), stoniness, st<strong>and</strong> age <strong>and</strong> TECI whereas<br />

vegetation class 4 showed the opposite pattern (Fig. 2b).<br />

<strong>Forest</strong> <strong>L<strong>and</strong>scapes</strong> <strong>and</strong> <strong>Global</strong> <strong>Change</strong>-New Frontiers in Management, Conservation <strong>and</strong> Restoration. Proceedings of the IUFRO L<strong>and</strong>scape Ecology<br />

Working Group International Conference, September 21-27, 2010, Bragança, Portugal. J.C. Azevedo, M. Feliciano, J. Castro & M.A. Pinto (eds.)<br />

2010, Instituto Politécnico de Bragança, Bragança, Portugal.

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