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Landscapes Forest and Global Change - ESA - Escola Superior ...

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N. Zurbriggen et al. 2010. Modeling feedbacks between avalanches <strong>and</strong> forests under a changing environment<br />

174<br />

Furthermore, to be able to model avalanches in sufficient detail, the cell size will be reduced<br />

from 100m side length to 25m. To ensure applicability of the forest-l<strong>and</strong>scape model with<br />

reduced cell size, we will compare model runs with both cell side lengths.<br />

The feedback effects will emerge once the avalanche release probability, flow dynamics,<br />

destruction, <strong>and</strong> regeneration subroutines are set up. These will have to be fine-tuned for current<br />

climatic <strong>and</strong> l<strong>and</strong>-use conditions to make sure that there are no runaway effects, which could<br />

develop due to the positive feedback cycle between forests <strong>and</strong> avalanches. Keeping the<br />

environmental conditions constant, we will make sure there is a balance between avalanches <strong>and</strong><br />

forest dynamics, by tuning the avalanche <strong>and</strong> forest parameters involved. Once the feedbacks<br />

are established, we will compare the output to current treeline positions <strong>and</strong> patterns, to estimate<br />

the precision of the model.<br />

In a sensitivity analysis, environmental conditions are again kept constant while disturbance<br />

intensity (frequency <strong>and</strong> magnitude) are varied, <strong>and</strong> the influence on output variables such as<br />

total biomass per area, location <strong>and</strong> pattern of treelines, <strong>and</strong> species composition will be<br />

analyzed. The merged model TreeMig-Av will then be applied to the study area of the Davos<br />

municipality, in the eastern Swiss Alps. The model will be validated against historical avalanche<br />

frequency data <strong>and</strong> compared to the output of other models such as those used for avalanche risk<br />

mapping. IPCC AR4 climatic change scenarios <strong>and</strong> different l<strong>and</strong> use scenarios will then be<br />

used to simulate avalanche release <strong>and</strong> forest cover scenarios for the next centuries, <strong>and</strong> to<br />

analyze how the feedback effects develop under the changing environmental conditions.<br />

3. Preliminary <strong>and</strong> expected results<br />

In the GLM, the final variables used for the estimation of avalanche release inside forests are<br />

similar between the two forest types: While the coniferous forest model equation uses slope<br />

angle, crown projection, <strong>and</strong> maximum gap size, the broadleaf forest equation uses slope angle,<br />

crown projection, <strong>and</strong> proportion of coniferous trees instead of the gap size.<br />

A sensitivity analysis in the current version of TreeMig, with single cell disturbances, showed<br />

that it is sensitive to changes in disturbance frequency <strong>and</strong> magnitude. <strong>Change</strong>s in these<br />

variables led to changes not only in total biomass, but also species composition <strong>and</strong> structural<br />

diversity in the affected cells. Furthermore, the single cell disturbance regime was compared to<br />

spatially connected disturbances on a transect, which confirmed the expected difference in<br />

regeneration patterns after the two disturbance types. The main feature of the spatially<br />

connected disturbance type is the potentially larger distance of a recently disturbed cell to the<br />

nearest vegetated cell, from where seeds may disperse into the disturbed cell. Regeneration<br />

dynamics therefore strongly depend on accurate simulation of dispersal kernels <strong>and</strong> size of the<br />

disturbed area.<br />

The Intermediate Disturbance Hypothesis (IDH) was shown to apply to TreeMig's disturbance<br />

rates at the single cell level, <strong>and</strong> for connected disturbances along the one-dimensional transect,<br />

<strong>and</strong> is expected to also apply to spatially connected disturbances in two-dimensional space.<br />

Highest species diversity was found at intermediate disturbance frequency <strong>and</strong> magnitude. The<br />

exact location of the peak diversity along the two axes, however, was influenced by altitude, i.e.<br />

by limitations given by climatic variables. For example, where growth <strong>and</strong> survival is limited by<br />

climate, a relatively high disturbance frequency could lead to the disappearance of the forest<br />

altogether. Here, the peak of diversity would shift to lower values of disturbance frequency. The<br />

IDH applies to species composition but not necessarily to forest structure (size distribution), as<br />

avalanches can increase the number of smaller trees relative to the larger trees. Applicability of<br />

IDH to structural diversity will further be studied using the adapted version of TreeMig.<br />

<strong>Forest</strong> <strong>L<strong>and</strong>scapes</strong> <strong>and</strong> <strong>Global</strong> <strong>Change</strong>-New Frontiers in Management, Conservation <strong>and</strong> Restoration. Proceedings of the IUFRO L<strong>and</strong>scape Ecology<br />

Working Group International Conference, September 21-27, 2010, Bragança, Portugal. J.C. Azevedo, M. Feliciano, J. Castro & M.A. Pinto (eds.)<br />

2010, Instituto Politécnico de Bragança, Bragança, Portugal.

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