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WHO monographs on selected medicinal plants - travolekar.ru

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<str<strong>on</strong>g>WHO</str<strong>on</strong>g> <str<strong>on</strong>g>m<strong>on</strong>ographs</str<strong>on</strong>g> <strong>on</strong> <strong>selected</strong> <strong>medicinal</strong> <strong>plants</strong><br />

odes, and inferior ovary of various shapes. The gourd-f<strong>ru</strong>it varies in size<br />

(15–40 cm in diameter) and shape in the many cultivated varieties, and the<br />

toughened, furrowed peduncle does not enlarge near it (4, 10).<br />

Plant material of interest: dried seeds<br />

General appearance<br />

The seeds are ovate, c<strong>on</strong>stricted at <strong>on</strong>e end forming a short, blunt extensi<strong>on</strong>;<br />

flat or weakly bic<strong>on</strong>vex; up to 25 mm l<strong>on</strong>g and 8–14 mm wide,<br />

3–4 mm thick; <strong>on</strong> both faces, close to the edge, is an encircling ridge and<br />

groove, 1–2 mm wide, absent from projecti<strong>on</strong>; testa creamy-white to pale<br />

beige with a satiny sheen, smooth or with irregular wrinkles; texture brittle,<br />

somewhat papery; inner surface of seed coat fawnish-white, dull,<br />

rough or scurfy. The seed is n<strong>on</strong>-endospermic. Embryo easily separated<br />

from testa, more or less entirely covered in a dark olive-green pellicle,<br />

with metallic lustre; light patches of inner seed coat may be adherent.<br />

Embryo pale greenish-yellow, oily; large, almost flat cotyled<strong>on</strong>s, small<br />

c<strong>on</strong>ical radical at c<strong>on</strong>stricted end of seed; inner surfaces of cotyled<strong>on</strong>s<br />

with three or five <strong>ru</strong>dimentary veins, palmately arranged (11).<br />

Organoleptic properties<br />

Odour: indistinct; taste: bland, oily and slightly nut-like (2, 10).<br />

Microscopic characteristics<br />

Epidermal cells of testa erect, prismatic, up to 200 μm l<strong>on</strong>g; walls thin,<br />

bearing slender vertical strips of thickening, usually sinuous in upper porti<strong>on</strong>;<br />

in surface view polyg<strong>on</strong>al, large with c<strong>on</strong>spicuous beads; starch<br />

grains abundant, up to 5 μm, simple but frequently clumped; a band,<br />

about six cells deep, of small, thin-walled, isodiametric or small, el<strong>on</strong>gated<br />

parenchymatous cells, finely reticulately thickened and str<strong>on</strong>gly lignified;<br />

a few larger, irregular simple pits; a single layer of large, sub-rectangular<br />

sclereids, lumen narrow, ovoid, walls very thick and c<strong>on</strong>spicuously layered,<br />

pits few and not well-defined, <strong>on</strong>ly middle lamella and primary wall<br />

str<strong>on</strong>gly lignified; in surface view the sclereids are somewhat el<strong>on</strong>gated<br />

and the anticlinal walls deeply sinuous. Internal to the sclereid band several<br />

layers of progressively larger lignified parenchymatous cells with<br />

very fine reticulate thickening; the cells, having short arm-like projecti<strong>on</strong>s,<br />

form a sp<strong>on</strong>gy, lacunose tissue; areas of c<strong>on</strong>tact between branches<br />

of cells have quite large simple perforati<strong>on</strong>s. Innermost layers less welldefined,<br />

parenchymatous, largest cells internally; greenish chromoplasts<br />

present. Cotyled<strong>on</strong> cells variable, very thin-walled, c<strong>on</strong>taining oily globules<br />

and aleur<strong>on</strong>e grains up to 4 μm in diameter (2).<br />

84

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