World Congress of Malacology Antwerp ... - Unitas Malacologica
World Congress of Malacology Antwerp ... - Unitas Malacologica World Congress of Malacology Antwerp ... - Unitas Malacologica
Three new species of minute heterobranchs (Gastropoda: Heterobranchia: Omalogyridae and Rissoellidae) from Namibia Rolán, Emilio 1 ; Luque, Ángel A. 2 ; Peñas, Anselmo 3 1. Museo de Historia Natural, Universidad de Santiago de Compostela, Santiago de Compostela, Spain, Email: erolan@emiliorolan.com 2. Laboratorio de Biología Marina, Departamento de Biología, Universidad Autónoma, C/ Darwin, 2, 28049 Madrid, Spain, Email: angel.luque@uam.es 3. Olérdola, 39, 5º C, 08800 Vilanova i la Geltrú, Barcelona, Spain, Email: anspm1@yahoo.es Namibian molluscan fauna is poorly known, with only about 140 species of marine molluscs recorded up to now. The superfamilies Rissoelloidea and Omalogyroidea, currently included in Heterobranchia, contain only the nominotypical families (Rissoellidae and Omalogyridae, respectively), and comprise minute marine gastropods which are still scarcely known all over the world. About 26 species of omalogyrids and 20 rissoellids have been described from the Atlantic, but there are no records of species of these two families from Namibia. Samples collected near Swakopmund (central Namibia) by washing algae from intertidal rocky pools result in several living specimens of two Omalogyridae and one Rissoellidae new species, which are here described. Retrotortina n. sp. differs from the only other known Atlantic species of the genus (Retrotortina fuscata Chaster, 1896, from the Mediterranean and NE. Atlantic) in having larger shell (up to 0.9 mm), less prominent abapical part, somewhat concave columellar edge and proportionally smaller aperture. Radula, operculum and head-foot characters of this new species are described. Ammonicera n. sp. (up to 0.67 mm) differs from the 17 known Atlantic species of this genus by its teleoconch lacking axial sculpture, with only a faint spiral cord at both upper part and basis. Rissoella n. sp. (up to 1.7 mm in length) differs from other congeneric species by shell and head-foot characters; the radula and operculum are also described. Towards a global taxonomic authority list for the Mollusca Rosenberg, Gary Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA, Email: rosenberg@ansp.org Much of systematist’s work is rummaging through the “graveyard” of synonymized and forgotten names to determine whether a candidate new taxon is indeed undescribed. Therefore an authoritative list of known taxa is essential to inventorying the world’s molluscan fauna. The authority list should minimally include all the names by which each taxon is known and would enable linking data from disparate sources, such as online databases of museum collections. Despite the recognized potential for global taxonomic authority lists to accelerate discovery and description of biodiversity, and the numerous programs and agencies calling for development of such lists, progress on molluscan databases has slowed. There are several successful regional projects: CLEMAM and CLECOM for Europe, Malacolog for the Western Atlantic, SOMBASE for the Southern Ocean, the Indo-Pacific marine database, but they are not yet interlinked. There are several impediments to going global. It is easier to get funding for regional projects than global ones, and to attract funding one must have discrete, visible products that provide services needed by the target audience. Each project thus has slightly different goals and practices. Various standards exist for required elements in taxonomic databases, and complying with all simultaneously is burdensome. Furthermore, scientists resist authority lists as impinging on taxonomic freedom. To progress toward a global list, we must recognize that it is necessary to have a federation of regional projects that feed into a central mechanism that supports multiple classifications. Editorial 186
structures are needed that allow maintenance by taxonomic experts while allowing community input, perhaps in Wiki format. The recently announced Encyclopedia of Life, which aims to create webpages for all known species, might help in developing such infrastructure. Sex and systematics in the Streptaxidae (Pulmonata: Stylommatophora) Rowson, Ben Dept. Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF10 3NP, Email: ben.rowson@museumwales.ac.uk Recently, sexual selection (often via sexual conflict) has been implicated in the diversification of organisms. The Streptaxidae are a little-studied, widely distributed and highly diverse family of carnivorous tropical land-snails. New data from an ongoing systematic study suggests sexual selection may have been a major force in the diversification of certain clades. A synapomorphy of the family is possession of a simple penis studded with sharp conchiolinous hooks. In different major African lineages this has been modified as follows: (A) taxa having serially repeating hooks, lacking spermatophores, and having a muscular vagina; (B) taxa having serially repeating hooks and penial stimulators containing novel, conchiolinous dart-like structures, lacking spermatophores, and having a muscular vagina; (C) taxa having differentiated, specialised hooks and configurations of penial pilasters, possibly functioning as stimulators, and possessing calc sacs, stylophores or spermatophores, but lacking a vagina. Type (A) and (B) taxa are moderately speciose, usually occurring in allopatry, while type (C) taxa are extremely speciose, often occurring in sympatry. These divisions are largely congruent with phylogenetic reconstructions based on non-genital morphology, mitochondrial (16S) and nuclear (actin exon) DNA. The latter also suggests that the majority of type (C) taxa are the product of a single, recent radiation. I interpret the genital differences between the three types as evidence of differences in the type and strength of sexual selection, and interpret the differences in diversity as evidence of the resulting response. Under this scenario, the genital features unique to type (C) taxa are seen as adaptations to rampant sexual conflict, and may also be key innovations that have promoted the recent diversification. The evidence for an alternative explanation, that of adaptive radiation relating to carnivory, is considered strong only for type (A) taxa. Improved phylogenetic reconstructions will allow a more robust test of these predictions in a comparative framework. Streptaxomorph shells: an evaluation and possible explanation Rowson, Ben Dept. Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF10 3NP, Email: ben.rowson@museumwales.ac.uk A “streptaxomorph” shell form, in which the coiling axis deviates substantially to the right during growth, is common among the Streptaxidae (Pulmonata: Stylommatophora). It does not occur in any plausible outgroup. This raises two questions: i) why does streptaxomorphy occur at all? and ii) has it arisen many times, or been repeatedly lost after being inherited by all streptaxids? This poster presents methods for describing the phenomenon in morphometric terms, offers explanations for its occurrence, and suggests a possible scenario for its evolution. Preliminary data indicate that the vast majority of streptaxid species are in fact axially deviated, either to the right (i.e., streptaxomorphy) or to the left (as in columnar or barrel-shaped species). Streptaxomorph shells are less frequent, but occupy a greater region of morphospace than left-deviated ones, including a region exploited by few other land snails (the “Cain gap” where height is approximately equal to diameter). However, alternative and more appropriate measurements indicate that this is misleading, with streptaxomorph 187
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Three new species <strong>of</strong> minute heterobranchs (Gastropoda: Heterobranchia: Omalogyridae and<br />
Rissoellidae) from Namibia<br />
Rolán, Emilio 1 ; Luque, Ángel A. 2 ; Peñas, Anselmo 3<br />
1. Museo de Historia Natural, Universidad de Santiago de Compostela, Santiago de Compostela,<br />
Spain,<br />
Email: erolan@emiliorolan.com<br />
2. Laboratorio de Biología Marina, Departamento de Biología, Universidad Autónoma, C/ Darwin, 2,<br />
28049 Madrid, Spain,<br />
Email: angel.luque@uam.es<br />
3. Olérdola, 39, 5º C, 08800 Vilanova i la Geltrú, Barcelona, Spain,<br />
Email: anspm1@yahoo.es<br />
Namibian molluscan fauna is poorly known, with only about 140 species <strong>of</strong> marine molluscs<br />
recorded up to now. The superfamilies Rissoelloidea and Omalogyroidea, currently included in<br />
Heterobranchia, contain only the nominotypical families (Rissoellidae and Omalogyridae,<br />
respectively), and comprise minute marine gastropods which are still scarcely known all over the<br />
world. About 26 species <strong>of</strong> omalogyrids and 20 rissoellids have been described from the Atlantic, but<br />
there are no records <strong>of</strong> species <strong>of</strong> these two families from Namibia.<br />
Samples collected near Swakopmund (central Namibia) by washing algae from intertidal rocky pools<br />
result in several living specimens <strong>of</strong> two Omalogyridae and one Rissoellidae new species, which are<br />
here described.<br />
Retrotortina n. sp. differs from the only other known Atlantic species <strong>of</strong> the genus (Retrotortina<br />
fuscata Chaster, 1896, from the Mediterranean and NE. Atlantic) in having larger shell (up to 0.9<br />
mm), less prominent abapical part, somewhat concave columellar edge and proportionally smaller<br />
aperture. Radula, operculum and head-foot characters <strong>of</strong> this new species are described.<br />
Ammonicera n. sp. (up to 0.67 mm) differs from the 17 known Atlantic species <strong>of</strong> this genus by its<br />
teleoconch lacking axial sculpture, with only a faint spiral cord at both upper part and basis.<br />
Rissoella n. sp. (up to 1.7 mm in length) differs from other congeneric species by shell and head-foot<br />
characters; the radula and operculum are also described.<br />
Towards a global taxonomic authority list for the Mollusca<br />
Rosenberg, Gary<br />
Academy <strong>of</strong> Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA,<br />
Email: rosenberg@ansp.org<br />
Much <strong>of</strong> systematist’s work is rummaging through the “graveyard” <strong>of</strong> synonymized and forgotten<br />
names to determine whether a candidate new taxon is indeed undescribed. Therefore an authoritative<br />
list <strong>of</strong> known taxa is essential to inventorying the world’s molluscan fauna. The authority list should<br />
minimally include all the names by which each taxon is known and would enable linking data from<br />
disparate sources, such as online databases <strong>of</strong> museum collections.<br />
Despite the recognized potential for global taxonomic authority lists to accelerate discovery and<br />
description <strong>of</strong> biodiversity, and the numerous programs and agencies calling for development <strong>of</strong> such<br />
lists, progress on molluscan databases has slowed. There are several successful regional projects:<br />
CLEMAM and CLECOM for Europe, Malacolog for the Western Atlantic, SOMBASE for the<br />
Southern Ocean, the Indo-Pacific marine database, but they are not yet interlinked.<br />
There are several impediments to going global. It is easier to get funding for regional projects than<br />
global ones, and to attract funding one must have discrete, visible products that provide services<br />
needed by the target audience. Each project thus has slightly different goals and practices. Various<br />
standards exist for required elements in taxonomic databases, and complying with all simultaneously<br />
is burdensome. Furthermore, scientists resist authority lists as impinging on taxonomic freedom.<br />
To progress toward a global list, we must recognize that it is necessary to have a federation <strong>of</strong><br />
regional projects that feed into a central mechanism that supports multiple classifications. Editorial<br />
186