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World Congress of Malacology Antwerp ... - Unitas Malacologica

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was also used as arena by the Romans. The city was famous with the statues. The statues signed by<br />

sculptures <strong>of</strong> Afrodisyas were found out in Italy, Greece and the other countries around<br />

Mediterranean. Also the marble blocks were exported. The marbles were obtained from the mine<br />

which still exists and is 2 kilometers east <strong>of</strong> Babadağ. The city was conquered by Turks at the 12th<br />

century A. D.<br />

The land snails were collected from the antique city and its surroundings with the permision <strong>of</strong> the<br />

local authority in April-2007.<br />

9 species were reported. The distribution <strong>of</strong> “Caracollina lenticula (Michaud, 1831)” as a<br />

synanthropic beach species in this antique settlement is discussed. Also the anatomical dissection <strong>of</strong><br />

the Helix sp. is given.<br />

We would like to thank for Ayşe Yıldırım for her helps in collecting the snails.<br />

Analysing speciation patterns – lessons from rissooidean and hydrocenid gastropods calling for<br />

the integration <strong>of</strong> methodologies<br />

Haase, Martin<br />

Vogelwarte, Zoologisches Institut und Museum der Universität Greifswald, Soldmannstraße 16, D-<br />

17489 Greifswald, Germany,<br />

Email: martin.haase@uni-greifswald.de<br />

Many small species <strong>of</strong> gastropod, especially those occurring in fresh water, have a low potential for<br />

dispersal and consequently restricted ranges. Yet, their diversity may be very high suggesting high<br />

rates <strong>of</strong> speciation. Small species tend to be poor in morphological characters resulting in a high<br />

degree <strong>of</strong> homoplasy. Often, species cannot be discriminated based on morphological data alone,<br />

because the taxonomic value <strong>of</strong> a morphological difference is difficult to interpret. Phylogenetic<br />

analyses based on morphological data are practically impossible. Genetic methods, especially<br />

sequence analyses, have brought enormous progress. However, there are a number <strong>of</strong> examples also<br />

<strong>of</strong> genetically cryptic species. Presenting cases from rissooidean and hydrocenid gastropods I will<br />

show that only the synthesis <strong>of</strong> morphological and genetic methods can really enhance taxonomic and<br />

phylogenetic analyses. In addition, the integrative approach is setting the stage for evolutionary<br />

analyses such as that <strong>of</strong> speciation patterns.<br />

Dynamic gastropods: morphological and genetic differentiation <strong>of</strong> the land snail Arianta<br />

arbustorum in an Alpine massif<br />

Haase, Martin 1 ; Mis<strong>of</strong>, Bernhard 2<br />

1. Vogelwarte, Zoologisches Institut und Museum der Universität Greifswald, Soldmannstraße 16, D-<br />

17489 Greifswald, Germany,<br />

Email: martin.haase@uni-greifswald.de<br />

2. Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D-53113 Bonn,<br />

Germany, Email: b.mis<strong>of</strong>.zfmk@uni-bonn.de<br />

Based on a sequence fragment <strong>of</strong> mtDNA and a very variable microsatellite locus we analyzed the<br />

genetic differentiation <strong>of</strong> more than 400 individuals <strong>of</strong> the land snail Arianta arbustorum collected in<br />

62 localities <strong>of</strong> the Alpine massif Gesäuse, where four morphotypes, characterized by geometric<br />

morphometrics, occur in close vicinity. We aimed at understanding actual and historical evolutionary<br />

processes among the morphotypes. With 135 haplotypes diverging up to 12.5% and 18 microsatellite<br />

alleles, genetic diversity was extremely high. The most common haplotype was found in all four<br />

morphotypes across the entire study area. Morph-specific haplotypes or alleles could not be<br />

identified. Phylogenetic analyses indicated that A. arbustorum has colonized the Gesäuse several<br />

times in consecutive waves and that introgression <strong>of</strong> neutral or near neutral markers across<br />

morphotypes and selection on shell shape, whose differentiation is apparently ancient, have played an<br />

important role. Gene flow was more likely between locally close, different morphotypes than among<br />

88

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