World Congress of Malacology Antwerp ... - Unitas Malacologica
World Congress of Malacology Antwerp ... - Unitas Malacologica World Congress of Malacology Antwerp ... - Unitas Malacologica
than 565 µ belongs to P. muscorum. The results could be applied for the Weichselian Pupilla shells from the loess series of the analysed region. Extreme variability in the radula of Anatomidae (Gastropoda: Vetigastropoda) Geiger, Daniel L. 1 ; Sasaki, Takenori 2 1. Santa Barbara Museum of Natural History – Invertebrate Zoology, 2559 Puesta del Sol Road, Santa Barbara, CA 93105, USA, Email: geiger@vetigastropoda.com 2. The University Museum, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan, Email: sasaki@um.u-tokyo.ac.jp Anatomidae is a family of small to moderate sized marine snails (1-11 mm) occurring world-wide from the shelf onwards to abyssal depth. Relatively few radulae have been investigated of only a handful of species. We have expanded the sampling significantly and have uncovered very different radular types to a degree usually associated with family-level differentiation. The variability observed to date by far exceeds that seen in Scissurellidae, for which more species have been examined. The shells of those species do not show any co-variation to the highly modified radulae. We have considered a number of possible factors that could explain those novel radula morphologies: - Depth: the novel radula types are from species occurring between several hundreds to a few thousand meters, but other species from equal or even greater depth show the more typical anatomid radula. - Hydrothermal influence: a couple of species, Anatoma janetae Geiger, 2006 and an undescribed species were collected in areas of hydrothermal influence, while a third, undescribed species with a highly modified radula did not occur in such a habitat. - Ocean basin: All species with modified radulae are from the Indian and Pacific Oceans, though sampling of species in the Atlantic is not as dense, and many other species in the Indian or Pacific Ocean have a typical anatomid radula. - Sexual dimorphism: Some molluscs exhibit sexual dimorphic radulae. In the anatomid species examined thus far, in species for which multiple specimens have been examined, they show the same radula, including one undescribed species with highly modified radula. The probability of examining one gender only of one species and only the other gender in a second species are exceedingly low. The best explanation is that local adaptations affect the radular development. Limited gut content analysis did not indicate any notable feature. Devil in the detail: Scissurellid systematics in the making (Gastropoda: Vetigastropoda) Geiger, Daniel L. Santa Barbara Museum of Natural History – Invertebrate Zoology, 2559 Puesta del Sol Road, Santa Barbara, CA 93105, USA, Email: geiger@vetigastropoda.com Scissurellids is a group of small marine snails of world-wide distribution. Work leading to a monograph is under way. The advantages and disadvantages of various sources of material (museums, private collectors, dealers, personal collecting) are evaluated. The majority of the literature can be compiled within about five years; Sherborn missed some taxa. Working on a family perceived as difficult offers many opportunities for significant discoveries (new morphologies, placement and monophyly of group, geographic and diversity patterns). The tropical-polar diversity gradient is confirmed both at the genus as well as at the species level. Common species are over-named, mainly due to underestimated intraspecific variation including scanning electron microscopy (SEM) of single specimens per species. Ontogenetic variability and poorly preserved rare species contribute to the overnaming. Similarly, mistaken diagnostic characters 72
can lead to erroneous beta taxonomy. Supposedly widely distributed species can be shown to represent discrete species. Approximately 89 species remain to be named, most from the Indo- Malayan Archipelago and New Zealand. Half of all lots and specimens are from the Indo-Pacific, while sampling effort declines exponentially to 1000 m; thereafter the decline is much steeper. The top nine of fifty collections assessed contain approximately 83% of all specimens and lots. Recognition and verification of discrete species requires SEM, but in most cases identification of established species can be done with a stereomicroscope. Advances in SEM technology (variable pressure, environmental) allow type specimens to be imaged without coating. Automontage z-stack integration does not work well for 1 mm white shells; compound microscopes do not adequately resolve some necessary fine structure. Some type specimens have crumbled to dust due to inappropriate storage conditions. A significant problem is the uncertain location and/or existence of types. Restrictive loan requirements of some countries (Brazil) make de facto impossible to include material from that area. The evolution of small body size in Vetigastropoda Geiger, Daniel L. 1 ; Thacker, Christine E. 2 1. Santa Barbara Museum of Natural History – Invertebrate Zoology, 2559 Puesta del Sol Road, Santa Barbara, CA 93105, USA, Email: geiger@vetigastropoda.com 2. Natural History Museum of Los Angeles County – Ichthyology Section, 900 Exposition Blvd., Los Angeles, CA 90007, USA, Email: thacker@nhm.org Vetigastropoda includes a variety of lineages ranging in size from 0.6 to 300 mm, and inhabiting depths of 0 to 6000 m. We use molecular phylogenetics, doubling the taxon sampling of our previous 40-taxon study of Vetigastropoda (Geiger & Thacker, 2005), to investigate the evolution of size and ecology in the group. We assembled a matrix of more than 2600 base pairs, of which approximately 1000 positions were parsimony informative, composed of Histone 3, COI, and 18S rRNA. The expanded data matrix includes several new and interesting lineages such as two skeneids, one seguenziid, Gorgolepsis, Temnocinclis, Incisura, Sinezona and Sukashitrochus. We emphasize the need of voucher specimens of small species, which usually harbor taxonomic uncertainty, and illustrate how to process minute species to gain the maximum information from single specimens (DNA, shell, radula). We apply continuous character state reconstruction to address size change and vertical colonization of the ocean in Vetigastropoda; both variables exhibit changes over more than three orders of magnitude. Three dimensional reconstruction with Amira 4.1 of the gonad of a scissurellid is used to explain and predict the smallest size in Vetigastropoda, which agrees with the smallest known Vetigastropoda. Freshwater Hydrobiidae of Morocco (Mollusca: Gastropoda): Inventory and preliminary red list at regional level Ghamizi, Mohamed University Cadi Ayad, Muséum d’Histoire Natrurelle de Marrakech, Faculté des Sciences Semlalia, BP 2390 Marrakech, Morocco, Email: mghamizi@yahoo.fr Forty five species belonging to fourteen hydrobiid genera are recorded from the continental freshwater hydrosystems of Morocco. Six genera with twenty three species are strictly stygobiont and collected from wells and hyporheic water of the streams. All are micro molluscs of the Hydrobiidae family, including forms that are not yet formally described. The restricted area of their habitat, in the groundwater systems with less possibilities of the dispersion, provides radiation in forms and high endemism. 73
- Page 101 and 102: fertilization as the predominant ma
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- Page 117 and 118: soil type and geological age) for P
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- Page 135 and 136: A DNA-based phylogeny resolves dive
- Page 137 and 138: and I. rositai. Besides, their phyl
- Page 139 and 140: The allozyme data showed no sign of
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- Page 177 and 178: New techniques yield new insights o
- Page 179 and 180: Motivation to resist sex in a simul
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- Page 183 and 184: in Utah (e.g. Oxychilus alliarius,
- Page 185 and 186: A recent review of the biogeography
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can lead to erroneous beta taxonomy. Supposedly widely distributed species can be shown to<br />
represent discrete species. Approximately 89 species remain to be named, most from the Indo-<br />
Malayan Archipelago and New Zealand. Half <strong>of</strong> all lots and specimens are from the Indo-Pacific,<br />
while sampling effort declines exponentially to 1000 m; thereafter the decline is much steeper. The<br />
top nine <strong>of</strong> fifty collections assessed contain approximately 83% <strong>of</strong> all specimens and lots.<br />
Recognition and verification <strong>of</strong> discrete species requires SEM, but in most cases identification <strong>of</strong><br />
established species can be done with a stereomicroscope. Advances in SEM technology (variable<br />
pressure, environmental) allow type specimens to be imaged without coating. Automontage z-stack<br />
integration does not work well for 1 mm white shells; compound microscopes do not adequately<br />
resolve some necessary fine structure. Some type specimens have crumbled to dust due to<br />
inappropriate storage conditions. A significant problem is the uncertain location and/or existence <strong>of</strong><br />
types. Restrictive loan requirements <strong>of</strong> some countries (Brazil) make de facto impossible to include<br />
material from that area.<br />
The evolution <strong>of</strong> small body size in Vetigastropoda<br />
Geiger, Daniel L. 1 ; Thacker, Christine E. 2<br />
1. Santa Barbara Museum <strong>of</strong> Natural History – Invertebrate Zoology, 2559 Puesta del Sol Road,<br />
Santa Barbara, CA 93105, USA,<br />
Email: geiger@vetigastropoda.com<br />
2. Natural History Museum <strong>of</strong> Los Angeles County – Ichthyology Section, 900 Exposition Blvd., Los<br />
Angeles, CA 90007, USA,<br />
Email: thacker@nhm.org<br />
Vetigastropoda includes a variety <strong>of</strong> lineages ranging in size from 0.6 to 300 mm, and inhabiting<br />
depths <strong>of</strong> 0 to 6000 m. We use molecular phylogenetics, doubling the taxon sampling <strong>of</strong> our previous<br />
40-taxon study <strong>of</strong> Vetigastropoda (Geiger & Thacker, 2005), to investigate the evolution <strong>of</strong> size and<br />
ecology in the group. We assembled a matrix <strong>of</strong> more than 2600 base pairs, <strong>of</strong> which approximately<br />
1000 positions were parsimony informative, composed <strong>of</strong> Histone 3, COI, and 18S rRNA. The<br />
expanded data matrix includes several new and interesting lineages such as two skeneids, one<br />
seguenziid, Gorgolepsis, Temnocinclis, Incisura, Sinezona and Sukashitrochus. We emphasize the<br />
need <strong>of</strong> voucher specimens <strong>of</strong> small species, which usually harbor taxonomic uncertainty, and<br />
illustrate how to process minute species to gain the maximum information from single specimens<br />
(DNA, shell, radula). We apply continuous character state reconstruction to address size change and<br />
vertical colonization <strong>of</strong> the ocean in Vetigastropoda; both variables exhibit changes over more than<br />
three orders <strong>of</strong> magnitude. Three dimensional reconstruction with Amira 4.1 <strong>of</strong> the gonad <strong>of</strong> a<br />
scissurellid is used to explain and predict the smallest size in Vetigastropoda, which agrees with the<br />
smallest known Vetigastropoda.<br />
Freshwater Hydrobiidae <strong>of</strong> Morocco (Mollusca: Gastropoda):<br />
Inventory and preliminary red list at regional level<br />
Ghamizi, Mohamed<br />
University Cadi Ayad, Muséum d’Histoire Natrurelle de Marrakech, Faculté des Sciences Semlalia,<br />
BP 2390 Marrakech, Morocco,<br />
Email: mghamizi@yahoo.fr<br />
Forty five species belonging to fourteen hydrobiid genera are recorded from the continental<br />
freshwater hydrosystems <strong>of</strong> Morocco. Six genera with twenty three species are strictly stygobiont and<br />
collected from wells and hyporheic water <strong>of</strong> the streams. All are micro molluscs <strong>of</strong> the Hydrobiidae<br />
family, including forms that are not yet formally described. The restricted area <strong>of</strong> their habitat, in the<br />
groundwater systems with less possibilities <strong>of</strong> the dispersion, provides radiation in forms and high<br />
endemism.<br />
73