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World Congress of Malacology Antwerp ... - Unitas Malacologica

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On the contrary in the family Turridae the loss <strong>of</strong> radula and venom apparatus is a unique event,<br />

which was described, until now, only in one species, Zemacies excelsa Sysoev & Bouchet, 2001.<br />

Loss <strong>of</strong> the radula in this species was combined with appearance <strong>of</strong> the unique foregut structure, the<br />

‘pyriform gland’. Basing on the presence <strong>of</strong> this unusual character in Z. excelsa the new<br />

nominotypical subfamily Zemaciinae was erected. Conchologically Zemaciinae are very close to<br />

Cochlespirinae (Turridae).<br />

Investigations <strong>of</strong> foregut anatomy <strong>of</strong> two still unnamed species <strong>of</strong> Horaiclavus (Turridae,<br />

Crassispirinae) for the first time in the subfamily demonstrated the absence in <strong>of</strong> the radula, venom<br />

gland and proboscis, as well as the presence <strong>of</strong> peculiar foregut structure, generally resembling<br />

‘pyriform gland’ <strong>of</strong> Zemacies excelsa. Other species <strong>of</strong> Horaiclavus possess radula <strong>of</strong> typical<br />

‘crassispirine’ type. Anatomy <strong>of</strong> the foregut <strong>of</strong> new species is superficially extremely similar to that<br />

<strong>of</strong> Z. excelsa, although conchological characters clearly preclude placing them in the same subfamily.<br />

Thus previously considered unique foregut arrangement <strong>of</strong> Zemaciinae appeared at least one more<br />

time independently in the subfamily Crassispirinae and probably does not have high taxonomic value.<br />

This requires reconsideration <strong>of</strong> the taxonomic status <strong>of</strong> subfamily Zemaciinae, which should be<br />

considered as a junior synonym <strong>of</strong> Cochlespirinae. The possible mechanism <strong>of</strong> functioning <strong>of</strong> the<br />

‘pyriform gland’ is suggested and discussed.<br />

Molecular Morphology <strong>of</strong> 18S rDNA <strong>of</strong> Bivalvia<br />

Feistel, S.; Haszprunar, G. 2<br />

Zoologische Staatssammlung München, Münchhausenstr. 21, 81247 München, Deutschland,<br />

Email: susanne@feistel.org; haszi@zsm.mwn.de<br />

The phylogeny <strong>of</strong> bivalves (Mollusca) consisting <strong>of</strong> about 20.000 species is still uncertain.<br />

Conventional morphology and sequence analyses did not provide conclusive results beyond the four<br />

subtaxa Protobranchia, Pteriomorpha, Heterodonta and Schizodonta. This study is concentrated on<br />

the folding patterns <strong>of</strong> slowly evolving 18S rDNA molecules in expectation <strong>of</strong> finding “deep<br />

phylogenetic” information, i.e. pre-Cambrian or Cambrian structural signatures.<br />

For this analysis, sequences <strong>of</strong> pteriomorph 18S rDNA with lengths <strong>of</strong> about 1800 base pairs each,<br />

available from European Ribosomal RNA Database and GenBank, have been studied.<br />

With publicly available s<strong>of</strong>tware, 3D molecular structures were created from the primary sequences<br />

und subsequently mapped into 2D patterns. The 2D/3D structures are known to be more conservative<br />

than the primary structure (the linear base sequence). Their morphological characters were detected,<br />

quantified and homologized for the generation <strong>of</strong> a property matrix. A phylogram has been<br />

constructed by means <strong>of</strong> a parsimony method. The method is described in detail and preliminary<br />

results are discussed.<br />

Phylogeny and morphological variability in land snails: the Sicilian Marmorana (Pulmonata,<br />

Helicidae)<br />

Fiorentino, Viviana 1 ; Salomone, Nicola 2 ; Manganelli, Guiseppe 1 ; Giusti, Folco 1<br />

1 Dipartimento di Scienze Ambientali “G. Sarfatti”, Università di Siena, Via P. A. Mattioli 4, I-53100<br />

Siena, Italy,<br />

Email: vivianafiorentino@unisi.it; manganelli@unisi.it; giustif@unisi.it<br />

2 Dipartimento di Biologia Evolutiva, Via A. Moro 2, 53100 Siena, Italy,<br />

Email: salomone@unisi.it<br />

Land snails have long been recognised as suitable organisms for studying phenotypic differentiation<br />

and phylogeny in relation to geographical distribution. Morphological data (shell and anatomical<br />

biometry on different geographical scales) and partial sequences (COI, 16S) were used to test<br />

whether morphological patterns matched phylogeny in a diversified group <strong>of</strong> Sicilian rock-dwelling<br />

land snails <strong>of</strong> the genus Marmorana. The two sets <strong>of</strong> data were also used to check the taxonomic and<br />

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