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RA 00110.pdf - OAR@ICRISAT

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Table 5. Prediction equations for interpopulation improvement methods—Reciprocal Half-sib Selection and Reciprocal<br />

Full-sib Selection.<br />

Half-sib Selection (RHS)<br />

Reciprocal Full-sib Selection (RFS)<br />

Note: = genetic variance among A x B full-sibs = in AB population.<br />

=genetic variance among half-sibs.<br />

line from the other population. Thus, every entry has<br />

one elite line parent and therefore, is a potential<br />

commercial hybrid, assuming further inbreeding<br />

and selection in the selected S, lines. Comstock<br />

(1979) concludes that RHS using the counterpart<br />

populations as testers is superior to RIS using pure<br />

line testers for improving the population cross. The<br />

RIS scheme has an advantage from a line development<br />

point of view and has been used by some<br />

commercial breeders.<br />

The major advantages and disadvantages of RRS<br />

procedures are outlined in Table 6.<br />

Response to Selection<br />

This section concentrates primarily on a comparison<br />

of S, line per se with reciprocal full-sib selection<br />

(RFS) because considerable data on these methods<br />

is available and because of their expected contrasting<br />

responses. Line per se selection would be expected<br />

to perform best only if additive types of effects were<br />

involved while reciprocal selection schemes like<br />

RFS would be best able to capitalize on nonadditive<br />

(especially overdominance) effects.<br />

A very large and comprehensive comparison of<br />

Table 6. Advantages and Disadvantages of Reciprocal Recurrent Selection (RRS) Procedures.<br />

Advantages<br />

1. Dominance types of relationships are important in cross-fertilized crops, and basing selection on crossed families seems<br />

the most logical way to take advantage of this heterotic vigor.<br />

2. Some forms of RRS can be integrated into applied line-development programs.<br />

3. The ultimate selection limit may be highest for this type of selection because any type of genetic variation that can<br />

contribute is exploited, though it may take a long time to make use of epistatic combinations on a population basis.<br />

Disadvantages<br />

1. Requires considerable labor for hand pollinations, both in making up families for testing and for recombination.<br />

2. Expensive, requires many yield-trial plots (especially for RHS and RIS).<br />

3. As in most other family schemes (with exception of intra-population full-sib selection) requires 3 growing seasons per<br />

cycle.<br />

115

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