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What Makes a Tree a Tree? Tree Biology 101 - Up

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<strong>What</strong> <strong>Makes</strong> a <strong>Tree</strong> a <strong>Tree</strong><br />

<strong>Tree</strong> <strong>Biology</strong> <strong>101</strong><br />

• <strong>Tree</strong> - woody plants<br />

that are large in size<br />

and have a single main<br />

trunk.<br />

• <strong>Tree</strong> - long lived<br />

woody perennials that<br />

compartmentalize.


<strong>Tree</strong>s are different from<br />

other plants because…<br />

If conditions remain<br />

favorable, they can live<br />

for a very long time


Let’s look at<br />

a tree and<br />

how it might<br />

grow over<br />

time….here’s<br />

how many<br />

seedlings<br />

look shortly<br />

after they<br />

sprout…..


In a few<br />

years……


….at about 10<br />

years or so


…perhaps<br />

in 30 to 40<br />

years


<strong>Tree</strong>s are genetically “wired”<br />

to live for a long time…..This<br />

tree is over 130 Years Old...


The Wye Oak (a white oak) was estimated<br />

to be over 460 years old before it fell…


This is what<br />

a 1700 year<br />

old tree<br />

looks like


This one is over<br />

2,300 years old!


275 ft tall<br />

33 feet in<br />

diameter, that<br />

is 103 ft in<br />

circumference


How Old Is The Oldest Living Thing On Earth<br />

“Methuselah” A Bristlecone Pine<br />

4,777<br />

YEARS<br />

OLD!!!!


<strong>What</strong> <strong>Makes</strong> A <strong>Tree</strong> A <strong>Tree</strong><br />

• <strong>Tree</strong> <strong>Biology</strong> - the study of structure and<br />

function, and the relationship between<br />

them<br />

– Anatomy - the study of the component<br />

parts of the tree<br />

– Physiology - the study of the biological and<br />

chemical processes within these<br />

components


Basic <strong>Tree</strong> Anatomy<br />

• All living organisms are made up of<br />

cells, tissues and organs<br />

• New cells form from the division of<br />

existing cells<br />

• In trees, these areas of cell division are<br />

called meristems


<strong>Tree</strong> Anatomy-Cells and<br />

Tissues continued<br />

• Apical Meristems - primary meristems that<br />

produce cells that result in elongation of<br />

roots and shoots<br />

• Lateral Meristems - secondary meristems<br />

that produce cells that result in an increase<br />

in diameter<br />

– Cambium - a lateral meristem that is a thin,<br />

continuous sheath of cells that divide to produce<br />

the vascular system of the tree:<br />

• Xylem – woody tissue, movement of water<br />

• Phloem – movement of food


Leaves


<strong>Tree</strong> Anatomy – Deciduous vs<br />

Evergreens<br />

• Evergreen <strong>Tree</strong>s – hold leaves for<br />

more than 1 year<br />

• Deciduous <strong>Tree</strong>s – shed leaves every<br />

year<br />

– Forms an Abscission Zone (A Z)<br />

• The A Z is formed as a result of changes in<br />

cells and growth regulators as temperatures<br />

drop and nights get longer in the Fall<br />

• A Z forms at the base of a leaf stem or<br />

“petiole” every year


<strong>Tree</strong> Anatomy-Leaves<br />

• Making Food is the Primary Function of<br />

Leaves! (Photosynthesis)<br />

– CO2 + H2O + Light Energy = Carbohydrates (Sugars) + O2<br />

– Occurs in:<br />

• Chlorophyll– green photo-reactive<br />

pigment found in the chloroplasts<br />

• Chloroplasts – specialized organelles<br />

found in plant cells that are the site of<br />

photosynthesis


From micro.magnet.fsu.edu


From ISA Introduction to Arboriculture <strong>Tree</strong> <strong>Biology</strong> CD


Stems


<strong>Tree</strong> Anatomy – Woody Section<br />

Earlywood - xylem produced in Spring<br />

Latewood - xylem produced later in Summer<br />

Growth Ring – visible rings created by the<br />

combination of early and latewood<br />

Ray cells – live parenchyma cells that transport<br />

sugars and other compounds through the trunk<br />

radially


Physiology -<br />

Compartmentalization<br />

• CODIT- Compartmentalization Of Decay In<br />

<strong>Tree</strong>s<br />

-Wall 1-resists vertical spread of decayxylem<br />

plugging (weakest wall)<br />

-Wall 2-resists inward spread of decaycompact<br />

latewood cells<br />

-Wall 3-resists lateral spread - ray cells<br />

-Wall 4-resists outward spread into new<br />

wood that is formed –(strongest<br />

wall)


From ISA Introduction to Arboriculture<br />

<strong>Tree</strong> <strong>Biology</strong> CD


Xylem continued<br />

• Ring Porous Wood – form wide vessels early<br />

in the growing season and narrower ones<br />

later<br />

– Elms<br />

– Oaks<br />

– Ash<br />

• Diffuse Porous Wood – form uniform sized<br />

vessels throughout growing season<br />

– Maples<br />

– Sycamore<br />

– Populus


Diffuse<br />

Porous Wood<br />

Ring Porous<br />

Wood<br />

University of Toronto<br />

University of Texas


<strong>Tree</strong> Anatomy - Phloem<br />

• Phloem – made up mostly of living cells<br />

– Responsible for the transport of food within the<br />

tree (sugars = photsynthates)<br />

• Transport occurs along a gradient<br />

• Transport requires energy<br />

– Comprised of:<br />

• Sieve Cells in conifers<br />

• Sieve Tube Elements and Companion Cells in<br />

hardwoods<br />

– As a tree grows, old phloem is crushed and may be<br />

absorbed into the tree or incorporated into the bark


<strong>Tree</strong> Anatomy - Bark<br />

• Bark - outer covering of a tree’s<br />

branches and stems composed of<br />

non-functional phloem and corky<br />

tissue that:<br />

-moderates temperature<br />

-offers some defense against injury<br />

-reduces water loss<br />

• Lenticels - small openings in bark<br />

that allow for gas exchange


Lenticels in Cherry Bark


<strong>Tree</strong> Anatomy - Stems<br />

• Hierarchy of stem composition:<br />

– Trunks support branches<br />

– Branches support other branches or twigs<br />

– Twigs support leaves, flowers and fruit<br />

• Buds occur:<br />

– At the tip of each twig (terminal or apical buds)<br />

– Along the twig (lateral or axillary buds)<br />

– At the base of each leaf ( flower and leaf buds)<br />

– Just under the bark (adventitious buds)<br />

• Dormant until released<br />

• When released they give rise to epicormic shoots


<strong>Tree</strong> Anatomy-Branches<br />

• Branches - are autonomous and usually<br />

produce and store enough carbohydrates to<br />

sustain themselves<br />

• Branch Collar-shoulder area where a branch<br />

joins another branch or trunk created by the<br />

overlapping xylem tissues<br />

– Strong attachment on bottom of branch to the parent stem<br />

– Weaker attachment on the top<br />

• Branch Bark Ridge-area of a branch union<br />

where the growth and development of the two<br />

adjoining limbs push the bark into a ridge<br />

• Included Bark-bark that is pushed inside a<br />

developing branch union, causing a<br />

weakened structure


From ISA Arborists’<br />

Certification Study<br />

Guide, Figure 8.2,<br />

1 st edition


From ISA Arborists’ Certification<br />

Study Guide, Figure 8.3, 1st edition<br />

From ISA<br />

Arborists’<br />

Certification<br />

Study Guide,<br />

Figure 8.5, 1 st<br />

edition


Roots


Mycorrhizae – the symbiotic association<br />

consisting of fungi and roots


Critical Root Zone


Physiology – Water Movement<br />

• Water is used for:<br />

– Maintaining cell turbidity<br />

– Transport of essential elements<br />

– Growth and Metabolism<br />

• Most of a trees water is lost through<br />

Transpiration<br />

• Pathway of water into roots = Osmosis:<br />

– Water goes from area of high water potential to<br />

area of low potential<br />

– Usually moves into roots from the soil, but can<br />

move out of roots into soil if potential is lower in<br />

the soil


<strong>Tree</strong> Physiology<br />

• Transpiration:<br />

– It is the loss of water vapor through the<br />

stomata<br />

– Cools leaves<br />

– Creates “transpirational pull”<br />

– Temperature, humidity & available water<br />

affect rates


The Cultivation<br />

of <strong>Tree</strong>s<br />

is the<br />

Cultivation of<br />

the Good…


…the<br />

Beautiful…


…and the Ennobling<br />

in Man…<br />

- J. Sterling Morton


QUESTIONS


WE NEED TO PLANT THE RIGHT<br />

TREE IN THE RIGHT PLACE!!!


We Cannot Control The<br />

Weather


<strong>Tree</strong> Physiology –<br />

The Vascular System<br />

• Phloem Transport:<br />

– Movement of photosynthate in the phloem<br />

• From areas of high sugar concentration to areas of lower sugar<br />

concentrations<br />

• Requires energy<br />

– Source - leaves, areas of high sugar concentration<br />

– Sink - plant parts that use more energy than they produce<br />

• Most plant parts can be sinks at some point in their life<br />

• Phloem transport occurs from leaves to roots, but most<br />

photosynthate is either used or stored in close proximity to where it<br />

is created<br />

• Radial Transport - horizontal movement of water and<br />

nutrients primarily through ray cells


From ISA<br />

Arborists<br />

Certification<br />

Study Guide,<br />

Figure 1.12, 1 st<br />

edition


Physiology – Growth Regulators<br />

• Growth & Development are results of the<br />

interaction of a tree’s genetic potential and<br />

the environmental conditions where it is<br />

growing<br />

– Plant Growth Regulators:<br />

• Naturally occurring substances that act to<br />

regulate plant growth and development<br />

• It only takes small quantities to affect growth<br />

• Classified as plant hormones:<br />

– Auxins<br />

– Cytokinins<br />

– Gibberellins<br />

– Abscisic acid<br />

– Ethylene


Physiology – Growth Regulators<br />

• Auxins<br />

– Produced primarily in shoot tips, but may be produced at other sites<br />

of cell division<br />

– Instrumental in root development<br />

– Involved with “tropisms”<br />

• Geotropism<br />

• Phototropism<br />

• Cytokinins<br />

– Produced primarily in the roots<br />

– Instrumental in shoot initiation and growth<br />

• Gibberellin<br />

– Growth regulator involved in cell elongation<br />

• Abscissic Acid<br />

– Plant growth substance that triggers leaf or fruit drop<br />

• Ethylene (gas)<br />

– Naturally occurring growth substance that triggers fruit ripening


Physiology – Growth Regulators<br />

• <strong>Tree</strong> form can be strongly directed by<br />

growth regulators<br />

– Decurrent tree form (rounded crown form)<br />

• Result of new lateral shoots outgrowing the original<br />

terminal shoot year after year<br />

• Indicates weak apical control<br />

– Excurrent tree form (upright crown form)<br />

• Results from strong apical control so that terminal shoot<br />

remains dominant as lateral shoots form<br />

• Most juvenile trees start out with excurrent form


Excurrent Habit<br />

Decurrent Habit<br />

from urbantree.org

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