Ecology of Red Maple Swamps in the Glaciated Northeast: A ...
Ecology of Red Maple Swamps in the Glaciated Northeast: A ... Ecology of Red Maple Swamps in the Glaciated Northeast: A ...
ences in nutrient levels, which were influenced by topographic position and hydrology. The most common type of red maple swamp encountered in the Damman and Kershner (1977) study was the Symplocarpus foetdus-Acer rubrum community that typically occurs in valley bottoms where soils are very poorly drained and fed by groundwater seepage (Fig. 4.1). These swamps are usually drained by a stream, so that surface water does not persist for long periods. If groundwater inflow is especially abundant and nutrient-rich, a Symplocapus-Acer rubrum- Ranunculus septentrionalis community is often found. Distinguishing species, besides swamp buttercup, in this floristically rich community include swamp saxifrage, bulbous bittercress, and golden ragwort. Upslope from the Symplocarpus- Acer rubrum community, in areas where soils are poorly drained but surface water is rarely present, a Betula alkghuniensis-Acer rubrum-Osmunda cinnamomea community is commonly found (Fig. 4.1).This transitional community frequently forms only a narrow belt at the bases of slopes; it is slightly drier and poorer in nutrients than the other two types of red maple forests. Pn devising a floristic classification for wetlands in the gneissic-schistose bedrock region of northwestern Connecticut, Messier (1980) also underscored the link between water regime and nutrient levels. He observed that, for a given nutrient regime, the type of wetland community was closely related to the elevation and degree of fluctuationof the water table. Figure 4.2 compares the extent of water level fluctuation during a single year among five red maple swamp communities and five other wetland types he encountered. In reviewing the following findings, remember that the extent of water level fluctuation may vary widely among years, even within the same swamp (Fig. 2.7). The Osmunda cinnarnomeu-Acer swamp occurred on peat soils of the valley floor, unlike the sloping sites described by Damman and Kershner (1977), and had a saturated water regime. The water table remained within 10-15 cm of the surface throughout the growing season, but surface water was present only briefly. The Rhododendron viscosum-Acer community occurred both in valley basins, where groundwater inflow was presumed to occur, and in basins farther upslope, which were perched above the local groundwater table. Water Quercus prinus - rubra Quercus !11ofoba and emsed bedr~ck I Fraxlnus Carya Fraxtnus-Acer saccharu Befula-Acer rubrum Fig. 4.1. Topsequences of plant communities on a till-covered gneiss hill in western Connecticut (after Damman and Kershner 1977). Left side of diagram represents normal topsequence; right side is that of certain south-facing slopes. Wetland communities are marked with an usterisk. Elevation of summit is between 350 and 400 m above sea level.
kvei fluctuation during the gmwing season was satuPated; by the end of the growing season, the comparable in the two Iucations, but the valley water table was commonly 60 crn or more below basina held much more water after spring snow- the surface. meit. Messier (1980) noted that varianfx of this paratley and Fahey (1986) identified three macommunity, dominated by different shrub spcies, jor forested wetland communities in a mixed conicould h di~tineished by the minimum pwing- fer--hardwood swamp in central New York hemseason wator level. Either RWPLdron visco- lock swamp; mixed conifer-red maple swamp, sum or Ilex uerl.ic&llntn appeared to be dominant larch phase; and mixed conifer-red maple swamp, where t,he? water t ~bk remained within 10 cm of whih pine phase. Using water level data gathered Lho surface (in 1978), while t7my:inium coryrnfw- wc?cMy during one growing season, the authors sum wits domirlar~t whore the water table dropped. denzomtrated that the distribution of woody speta at least 20 cm hilow the surface. cies was controlled largely by the mean depth h Tfle remaining tflree red maple swamp cornmunities, C~MX sbrictn - Awr, Ccirex lacustris-Acer, the water table during the growing season and the duration of the summer drawdown. Red maple arrd Symplor~irpus -Acer, were observed both in was the dominant tree in the severely flooded valley hotbms and in association with springs at sites, where the water level was highest and the the bases of valley . s~o~B. 1~1 valley bottoms, water period of drawdown was 8 weeks or less. Hemlock levels for all the commur~ities ranged from about swamp had a lower mean water level, but shorter 20 to $0 cxtr. aPK>vca C.c.3 20 to crrz below the surface drawdown period, than the mixed conifer-red maduring the growing season; howovc~r, during &larch, aurfa~e wt~f~e~r was eonsidcrribly deeper in Lha auctgt- to~xxrrlrrtlil ics (Fig. 4.2). At aftring siteer, 190th sedge rorr~nzurkitien tltd water levels within 5 -10 crn of ttie s~rl*f,rrcb througt~oul the growing aeasol.1 arrd were fl~wd~d to A ciopth of aniy 10- 213 c.rn during tlzc* elrring. ??re$ sedge cornrxrunities differed chicfly in xxtitrient ~Catue, the Caren Ictcustris-Acrr comrnunit~y wsccurrirlg in slightly ple communities (Table 4.1). Mean depth to the water table was also one of the key factors separatirrg four ground vegetation associations occurrirxg in the mixed conifer-red maple swamps; ash cont~nt and bulk density of the organic soils were other important factors @able 4.1). P~ratley and Fahey (1986) concluded that, in areas of the forested wetland with low mean water levels, the duration of summer drawdown was an ricizrr arms. The SS~~nrplwnr~~us- Arc~r coxn~nuniCy in:portant factor influencing both overstory and OCEUIT~*~~ at, ~prinh: sittas that wcw cttlly seaso~lally ground vegetation composition. Where mean Fig. 4.2 Water level fluctuation in red maple swamps and other wetland communities of northwestern Connecticut (redrafted from Messier 1980).
- Page 21 and 22: Fig. 1.4. The range of red maple (a
- Page 23 and 24: ~ b 1.5, h fircort~ of Wet landc Ir
- Page 25 and 26: Fig. 22. &~lrit~ve Irtndtici~pe yos
- Page 27 and 28: egional groundwakr table by the roc
- Page 29 and 30: y e~rr~mtrtm~~,irzst ion. Cat~t,inu
- Page 31 and 32: inflow^ Outflows OF SWQ SWI Fig. 25
- Page 33 and 34: Fig, 23.6,Soasonally flmded red map
- Page 35 and 36: Fig. 27. Water levels in six mode I
- Page 37 and 38: The duration of soil saturation has
- Page 39 and 40: Organic soils are always very poorl
- Page 41 and 42: Chapter 3. The Plant Community The
- Page 43 and 44: .dar02f 'B Xi? s%u?mma -?sway+xqq p
- Page 47 and 48: Community S tructare Red maple swam
- Page 49 and 50: suggesta a strong correlation betwe
- Page 51 and 52: Table 3.2. Stmtuml chumcteristics o
- Page 53 and 54: Table 3.3. Continued. "- -- - Speci
- Page 55 and 56: Table __ 3.3. _ Continued ._.lll.__
- Page 57 and 58: Table 3.3. Continued. . * - __.. ^.
- Page 59 and 60: Zone I II III TV C" Drppnmriad* sp.
- Page 61 and 62: on Long Island, pin oak, swamp whit
- Page 63 and 64: fern moea (?ki.c&inz &limfulum), an
- Page 65 and 66: countered (Table 3.3). The herb lay
- Page 67 and 68: ~akareous Seepage Swamps England si
- Page 69 and 70: and globeflower are. listed in four
- Page 71: was the most likely reason for diff
- Page 75 and 76: Table 4.2. .Rektit.e ~bmchnce w Q)
- Page 77 and 78: difficuit to delineate in many inst
- Page 79 and 80: in the librature btlx>~ithe moi~ttl
- Page 81 and 82: Origin. an$ Relationship to Water R
- Page 83 and 84: Influence on Swamp Vegetation Flori
- Page 85 and 86: taka wpra-x li)wcir, t raac* df~rla
- Page 87 and 88: dwuL 4.3 6.3. C ~ ~ ~ strisk- P C I
- Page 89 and 90: Chapter 5. Ecosystem Processes Irr
- Page 91 and 92: - 0 2 E E 0l *" C 8 g 00 - c_ m 3 -
- Page 93 and 94: Ehnzdoltf (IWj wm mmht~nt, rru@w on
- Page 95 and 96: 1976; I">irwia RE& vari iier Vdk 18
- Page 97 and 98: Xletritus Exprort and 'sod Chain Su
- Page 99: Chapter 6. Wetland Dynamics Most no
- Page 102 and 103: since forested wetland is the endpo
- Page 104 and 105: kettle bogs, lakes, or large rivers
- Page 106 and 107: mmdwakr depression wetlands. Becaus
- Page 108 and 109: (Mniotila varia), regularly breed i
- Page 110 and 111: EXusbtand arid Eddleman (1Y30) quan
- Page 112 and 113: census results. Twenty-five (40%) o
- Page 114 and 115: elated to avian richness and abunda
- Page 116 and 117: Fig. 7.6. Wood duck (Aix sponsa). T
- Page 118 and 119: Table 7.5. Small-mammal communities
- Page 120 and 121: ing the latter years of flowage occ
ences <strong>in</strong> nutrient levels, which were <strong>in</strong>fluenced by<br />
topographic position and hydrology.<br />
The most common type <strong>of</strong> red maple swamp<br />
encountered <strong>in</strong> <strong>the</strong> Damman and Kershner (1977)<br />
study was <strong>the</strong> Symplocarpus foetdus-Acer rubrum<br />
community that typically occurs <strong>in</strong> valley<br />
bottoms where soils are very poorly dra<strong>in</strong>ed and<br />
fed by groundwater seepage (Fig. 4.1). These<br />
swamps are usually dra<strong>in</strong>ed by a stream, so that<br />
surface water does not persist for long periods. If<br />
groundwater <strong>in</strong>flow is especially abundant and<br />
nutrient-rich, a Symplocapus-Acer rubrum-<br />
Ranunculus septentrionalis community is <strong>of</strong>ten<br />
found. Dist<strong>in</strong>guish<strong>in</strong>g species, besides swamp<br />
buttercup, <strong>in</strong> this floristically rich community <strong>in</strong>clude<br />
swamp saxifrage, bulbous bittercress, and<br />
golden ragwort. Upslope from <strong>the</strong> Symplocarpus-<br />
Acer rubrum community, <strong>in</strong> areas where soils are<br />
poorly dra<strong>in</strong>ed but surface water is rarely present,<br />
a Betula alkghuniensis-Acer rubrum-Osmunda<br />
c<strong>in</strong>namomea community is commonly found<br />
(Fig. 4.1).This transitional community frequently<br />
forms only a narrow belt at <strong>the</strong> bases <strong>of</strong> slopes; it<br />
is slightly drier and poorer <strong>in</strong> nutrients than <strong>the</strong><br />
o<strong>the</strong>r two types <strong>of</strong> red maple forests.<br />
Pn devis<strong>in</strong>g a floristic classification for wetlands<br />
<strong>in</strong> <strong>the</strong> gneissic-schistose bedrock region <strong>of</strong> northwestern<br />
Connecticut, Messier (1980) also underscored<br />
<strong>the</strong> l<strong>in</strong>k between water regime and nutrient<br />
levels. He observed that, for a given nutrient<br />
regime, <strong>the</strong> type <strong>of</strong> wetland community was<br />
closely related to <strong>the</strong> elevation and degree <strong>of</strong> fluctuation<strong>of</strong><br />
<strong>the</strong> water table. Figure 4.2 compares <strong>the</strong><br />
extent <strong>of</strong> water level fluctuation dur<strong>in</strong>g a s<strong>in</strong>gle<br />
year among five red maple swamp communities<br />
and five o<strong>the</strong>r wetland types he encountered. In<br />
review<strong>in</strong>g <strong>the</strong> follow<strong>in</strong>g f<strong>in</strong>d<strong>in</strong>gs, remember that<br />
<strong>the</strong> extent <strong>of</strong> water level fluctuation may vary<br />
widely among years, even with<strong>in</strong> <strong>the</strong> same swamp<br />
(Fig. 2.7).<br />
The Osmunda c<strong>in</strong>narnomeu-Acer swamp occurred<br />
on peat soils <strong>of</strong> <strong>the</strong> valley floor, unlike <strong>the</strong><br />
slop<strong>in</strong>g sites described by Damman and Kershner<br />
(1977), and had a saturated water regime. The<br />
water table rema<strong>in</strong>ed with<strong>in</strong> 10-15 cm <strong>of</strong> <strong>the</strong> surface<br />
throughout <strong>the</strong> grow<strong>in</strong>g season, but surface<br />
water was present only briefly. The Rhododendron<br />
viscosum-Acer community occurred both <strong>in</strong> valley<br />
bas<strong>in</strong>s, where groundwater <strong>in</strong>flow was presumed<br />
to occur, and <strong>in</strong> bas<strong>in</strong>s far<strong>the</strong>r upslope, which were<br />
perched above <strong>the</strong> local groundwater table. Water<br />
Quercus pr<strong>in</strong>us - rubra<br />
Quercus !11<strong>of</strong>oba<br />
and<br />
emsed bedr~ck<br />
I<br />
Fraxlnus Carya<br />
Fraxtnus-Acer saccharu<br />
Befula-Acer rubrum<br />
Fig. 4.1. Topsequences <strong>of</strong> plant communities on a till-covered gneiss hill <strong>in</strong> western Connecticut (after Damman<br />
and Kershner 1977). Left side <strong>of</strong> diagram represents normal topsequence; right side is that <strong>of</strong> certa<strong>in</strong><br />
south-fac<strong>in</strong>g slopes. Wetland communities are marked with an usterisk. Elevation <strong>of</strong> summit is between 350 and<br />
400 m above sea level.