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Ecology of Red Maple Swamps in the Glaciated Northeast: A ...

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Table 1.3. Pr<strong>in</strong>cipal tree species <strong>in</strong> upland and wetland forests <strong>of</strong> <strong>the</strong>glaciated Nor<strong>the</strong>ast, by forest region<br />

(based primarily on Lull 1968; names modi%d after Little 1979).<br />

Forest region<br />

Spruce-fir<br />

Beech-birchmaplen<br />

White p<strong>in</strong>ehendockhardwood<br />

Upland forests<br />

<strong>Red</strong> spruce<br />

White spruce<br />

Black spruce<br />

Balsam fi<br />

American beech<br />

Yellow birch<br />

Sugar maple<br />

American beech<br />

Yellow birch<br />

Sugar maple<br />

Eastern hemlock<br />

Black birch<br />

<strong>Red</strong> maple<br />

Basswood<br />

White ash<br />

Nor<strong>the</strong>rn red oak<br />

White p<strong>in</strong>e<br />

Eastern hemlock<br />

Nor<strong>the</strong>rn red oak<br />

Wetland forests<br />

Black spruce<br />

Tamarack<br />

Nor<strong>the</strong>rn white<br />

cedar<br />

Balsam fir<br />

<strong>Red</strong> maple<br />

Black ash<br />

Nor<strong>the</strong>rn white<br />

cedar<br />

Black spruce<br />

Tamarack<br />

<strong>Red</strong> maple<br />

Black ash<br />

nAlso frequently referred Lo us rlortliern hardwoods.<br />

/<br />

Pitch p<strong>in</strong>e-<br />

<strong>Red</strong> maple hardwood<br />

Ashes<br />

Eastern hemlock<br />

I<br />

I<br />

Forest region<br />

(cont<strong>in</strong>ued)<br />

Upland forests<br />

-- - --<br />

American beech<br />

Yellow birch<br />

Sugar maple<br />

O<strong>the</strong>r oaks<br />

Yellow-poplar<br />

Hickories<br />

<strong>Red</strong> maple<br />

White oak<br />

Nor<strong>the</strong>rn red oak<br />

Black oak<br />

Scarlet oak<br />

Chestnut oak<br />

Hickories<br />

Yellow-poplar<br />

Pitch p<strong>in</strong>e<br />

Bear oak<br />

Wetland forests<br />

White p<strong>in</strong>e<br />

Atlantic white<br />

cedar<br />

Fbd maple<br />

Atlantic white<br />

cedar<br />

Black gum<br />

<strong>Red</strong> maple<br />

Black g um<br />

Atlantic white<br />

cedar<br />

types occurr<strong>in</strong>g <strong>in</strong> <strong>the</strong> glaciated Nor<strong>the</strong>ast. Of <strong>the</strong><br />

five forest cover types <strong>in</strong> which it is a major component,<br />

three (white p<strong>in</strong>e-nor<strong>the</strong>rn red oak-red maple,<br />

gray birch-red maple, and black cherry-maple)<br />

are upland forest types, one (black ash-American<br />

elm-red maple) is a wetland type, and one (red<br />

maple) may occur on ei<strong>the</strong>r wetland or upland sites.<br />

So, while red maple is <strong>the</strong> dom<strong>in</strong>ant tree <strong>in</strong> <strong>the</strong> vast<br />

majority <strong>of</strong> broad-leaved deciduous wetland forests<br />

<strong>in</strong> <strong>the</strong> Nor<strong>the</strong>ast, it is classified as a facultative<br />

species, that is, one that occurs <strong>in</strong> wetlands from<br />

one-third to two-thirds <strong>of</strong> <strong>the</strong> time (Reed 1988).<br />

The distribution <strong>of</strong> red maple forested wetlands<br />

generally co<strong>in</strong>cides with <strong>the</strong> comb<strong>in</strong>ed distributions<br />

<strong>of</strong> <strong>the</strong> black ash-American elm-red maple cover<br />

type (SAF type no. 39) and <strong>the</strong> red maple type (no.<br />

108). The former type is found throughout <strong>the</strong> glaciated<br />

Nor<strong>the</strong>ast and <strong>the</strong> Great Lakes States, and<br />

from sou<strong>the</strong>rn Manitoba to Newfoundland (Eyre<br />

1980). In <strong>the</strong> Great Lakes States, black ash may be<br />

as abundant as elm and red maple <strong>in</strong> t.his cover<br />

type, but elsewhere it usually composes a small<br />

percentage <strong>of</strong> <strong>the</strong> stand. American elm has greatly<br />

decl<strong>in</strong>ed <strong>in</strong> abundance due tcr Dutch elm disease, so<br />

red maple has become <strong>the</strong> dom<strong>in</strong>ant species <strong>in</strong> <strong>the</strong> disturbed sites.<br />

black ash-American elm-red maple type throughout<br />

<strong>the</strong> Nor<strong>the</strong>ast.<br />

The red maple cover type (SAF no. 108) is most<br />

common <strong>in</strong> New England, <strong>the</strong> Middle Atlantic<br />

States, <strong>the</strong> Upper Pen<strong>in</strong>sula <strong>of</strong> Michigan, and<br />

nor<strong>the</strong>astern Wiscons<strong>in</strong>. Toward <strong>the</strong> western and<br />

sou<strong>the</strong>rn limits <strong>of</strong> its range, this type generally<br />

occurs on wetland soils; <strong>in</strong> New England and <strong>the</strong><br />

Upper Pen<strong>in</strong>sula <strong>of</strong> Michigan, it is found both <strong>in</strong><br />

wetlands and on dry, sandy, or rocky upland sites.<br />

In Pennsylvania, most red maple stands are found<br />

on mesic to dry upland sites (Eyre 1980).<br />

The SAF established <strong>the</strong> red maple forest cover<br />

type <strong>in</strong> 1988; before that, red maple was merely<br />

listed as a codom<strong>in</strong>ant or associated species <strong>in</strong> a<br />

number <strong>of</strong> o<strong>the</strong>r types. The dramatic <strong>in</strong>crease <strong>in</strong> <strong>the</strong><br />

proportion <strong>of</strong> red maple <strong>in</strong> many stands s<strong>in</strong>ce <strong>the</strong><br />

previous SAF classification (SAM' 1954) has been<br />

attributed to disturbances such as logg<strong>in</strong>g and fwe<br />

and <strong>the</strong> progressive elim<strong>in</strong>ation <strong>of</strong> American elm by<br />

Dutch elm disease (Eyre 1980). Production <strong>of</strong> heavy<br />

seed crops nearly every spr<strong>in</strong>g, rapid seed germ<strong>in</strong>ation,<br />

and vigorous sprout<strong>in</strong>g from stumps m d dmaged<br />

seedl<strong>in</strong>gs give red maple a competitive advantage<br />

over associated species on a wide variety <strong>of</strong>

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