Ecology of Red Maple Swamps in the Glaciated Northeast: A ...
Ecology of Red Maple Swamps in the Glaciated Northeast: A ...
Ecology of Red Maple Swamps in the Glaciated Northeast: A ...
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Table 1.3. Pr<strong>in</strong>cipal tree species <strong>in</strong> upland and wetland forests <strong>of</strong> <strong>the</strong>glaciated Nor<strong>the</strong>ast, by forest region<br />
(based primarily on Lull 1968; names modi%d after Little 1979).<br />
Forest region<br />
Spruce-fir<br />
Beech-birchmaplen<br />
White p<strong>in</strong>ehendockhardwood<br />
Upland forests<br />
<strong>Red</strong> spruce<br />
White spruce<br />
Black spruce<br />
Balsam fi<br />
American beech<br />
Yellow birch<br />
Sugar maple<br />
American beech<br />
Yellow birch<br />
Sugar maple<br />
Eastern hemlock<br />
Black birch<br />
<strong>Red</strong> maple<br />
Basswood<br />
White ash<br />
Nor<strong>the</strong>rn red oak<br />
White p<strong>in</strong>e<br />
Eastern hemlock<br />
Nor<strong>the</strong>rn red oak<br />
Wetland forests<br />
Black spruce<br />
Tamarack<br />
Nor<strong>the</strong>rn white<br />
cedar<br />
Balsam fir<br />
<strong>Red</strong> maple<br />
Black ash<br />
Nor<strong>the</strong>rn white<br />
cedar<br />
Black spruce<br />
Tamarack<br />
<strong>Red</strong> maple<br />
Black ash<br />
nAlso frequently referred Lo us rlortliern hardwoods.<br />
/<br />
Pitch p<strong>in</strong>e-<br />
<strong>Red</strong> maple hardwood<br />
Ashes<br />
Eastern hemlock<br />
I<br />
I<br />
Forest region<br />
(cont<strong>in</strong>ued)<br />
Upland forests<br />
-- - --<br />
American beech<br />
Yellow birch<br />
Sugar maple<br />
O<strong>the</strong>r oaks<br />
Yellow-poplar<br />
Hickories<br />
<strong>Red</strong> maple<br />
White oak<br />
Nor<strong>the</strong>rn red oak<br />
Black oak<br />
Scarlet oak<br />
Chestnut oak<br />
Hickories<br />
Yellow-poplar<br />
Pitch p<strong>in</strong>e<br />
Bear oak<br />
Wetland forests<br />
White p<strong>in</strong>e<br />
Atlantic white<br />
cedar<br />
Fbd maple<br />
Atlantic white<br />
cedar<br />
Black gum<br />
<strong>Red</strong> maple<br />
Black g um<br />
Atlantic white<br />
cedar<br />
types occurr<strong>in</strong>g <strong>in</strong> <strong>the</strong> glaciated Nor<strong>the</strong>ast. Of <strong>the</strong><br />
five forest cover types <strong>in</strong> which it is a major component,<br />
three (white p<strong>in</strong>e-nor<strong>the</strong>rn red oak-red maple,<br />
gray birch-red maple, and black cherry-maple)<br />
are upland forest types, one (black ash-American<br />
elm-red maple) is a wetland type, and one (red<br />
maple) may occur on ei<strong>the</strong>r wetland or upland sites.<br />
So, while red maple is <strong>the</strong> dom<strong>in</strong>ant tree <strong>in</strong> <strong>the</strong> vast<br />
majority <strong>of</strong> broad-leaved deciduous wetland forests<br />
<strong>in</strong> <strong>the</strong> Nor<strong>the</strong>ast, it is classified as a facultative<br />
species, that is, one that occurs <strong>in</strong> wetlands from<br />
one-third to two-thirds <strong>of</strong> <strong>the</strong> time (Reed 1988).<br />
The distribution <strong>of</strong> red maple forested wetlands<br />
generally co<strong>in</strong>cides with <strong>the</strong> comb<strong>in</strong>ed distributions<br />
<strong>of</strong> <strong>the</strong> black ash-American elm-red maple cover<br />
type (SAF type no. 39) and <strong>the</strong> red maple type (no.<br />
108). The former type is found throughout <strong>the</strong> glaciated<br />
Nor<strong>the</strong>ast and <strong>the</strong> Great Lakes States, and<br />
from sou<strong>the</strong>rn Manitoba to Newfoundland (Eyre<br />
1980). In <strong>the</strong> Great Lakes States, black ash may be<br />
as abundant as elm and red maple <strong>in</strong> t.his cover<br />
type, but elsewhere it usually composes a small<br />
percentage <strong>of</strong> <strong>the</strong> stand. American elm has greatly<br />
decl<strong>in</strong>ed <strong>in</strong> abundance due tcr Dutch elm disease, so<br />
red maple has become <strong>the</strong> dom<strong>in</strong>ant species <strong>in</strong> <strong>the</strong> disturbed sites.<br />
black ash-American elm-red maple type throughout<br />
<strong>the</strong> Nor<strong>the</strong>ast.<br />
The red maple cover type (SAF no. 108) is most<br />
common <strong>in</strong> New England, <strong>the</strong> Middle Atlantic<br />
States, <strong>the</strong> Upper Pen<strong>in</strong>sula <strong>of</strong> Michigan, and<br />
nor<strong>the</strong>astern Wiscons<strong>in</strong>. Toward <strong>the</strong> western and<br />
sou<strong>the</strong>rn limits <strong>of</strong> its range, this type generally<br />
occurs on wetland soils; <strong>in</strong> New England and <strong>the</strong><br />
Upper Pen<strong>in</strong>sula <strong>of</strong> Michigan, it is found both <strong>in</strong><br />
wetlands and on dry, sandy, or rocky upland sites.<br />
In Pennsylvania, most red maple stands are found<br />
on mesic to dry upland sites (Eyre 1980).<br />
The SAF established <strong>the</strong> red maple forest cover<br />
type <strong>in</strong> 1988; before that, red maple was merely<br />
listed as a codom<strong>in</strong>ant or associated species <strong>in</strong> a<br />
number <strong>of</strong> o<strong>the</strong>r types. The dramatic <strong>in</strong>crease <strong>in</strong> <strong>the</strong><br />
proportion <strong>of</strong> red maple <strong>in</strong> many stands s<strong>in</strong>ce <strong>the</strong><br />
previous SAF classification (SAM' 1954) has been<br />
attributed to disturbances such as logg<strong>in</strong>g and fwe<br />
and <strong>the</strong> progressive elim<strong>in</strong>ation <strong>of</strong> American elm by<br />
Dutch elm disease (Eyre 1980). Production <strong>of</strong> heavy<br />
seed crops nearly every spr<strong>in</strong>g, rapid seed germ<strong>in</strong>ation,<br />
and vigorous sprout<strong>in</strong>g from stumps m d dmaged<br />
seedl<strong>in</strong>gs give red maple a competitive advantage<br />
over associated species on a wide variety <strong>of</strong>