Ecology of Red Maple Swamps in the Glaciated Northeast: A ...

Ecology of Red Maple Swamps in the Glaciated Northeast: A ... Ecology of Red Maple Swamps in the Glaciated Northeast: A ...

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Seabard Lowlarad =@om $a more than 1 ,504) rn in the 'M7kite Mounts md Ahndacb. Coastal areas (incluw the Great Lakes region) generally are relatively flat> while mountainous regions are chcaracterized by step slopes and nmow valleys. The bulk of the Northeast falls within the New England Upland and Glaciated Allegheny Plateau regions, where moderate elevations (150-600 m), rolling hills, and nmw river valleys predominate. Bedrock types include primarily igneous and metamorphic mks through most of New England and in the Adkrondack Mountains and limestone, sandstone, and shale in much of the rest of the Northeast vable 1.1). Unstratified glacial deposits, more commonly known as till, predominate in the region. Stratified deposits are found in abundance in lowlands near the glacial limit, especially in southern New England (Seaboard Lowland) and northern New Jersey (Coastal Main and Piedmont), but also in deep preglacial valleys of central New Uork and in low-lying areas within the Great Lakes and St. Lawrence Valley physiographic regions. Marine sediments occur in parts of the New England Seaboard Lowland and St. Lawrence Val- Iey (Ferneman 1938; Lull 1968; Cunningham and Ciolkosz 1984). Climate G h t e in the Northeast is highly varied because of the wide range of physiographic conditions and the idu- of the Atlantic Ocean and Great Lakes (Cunnhgham and Ciolkcsz 1934). Variability in time and is probably the most mntpicuous as@ of the region's climate. There are wide ranges in daily and mual tempsmh, wide variations in temperature and pmipihtion for the same month or season in different years, and marked fluctuations in weather conditions over short periods (Ruffner 1985). bughoutthe glaciated Northeast, precipitation is evenly distxibuted over the year. Total annual precipitation ranges from more than 135 cm incertainamas of the White Mountains, Green Mountains, and Catskhllstolessthan75cmin~eGreatLakesregionand the Lake Champlain basin (Moody et it. 1986). Mean annual precipitation values for the variou9 northeastern stah are similar, however, gendy averaging 102- 122 cm. Total snowfall varies greatly overthe glaci- ated Northeast, Annual m10unts from less than 81cmonthe~astalRaintoasmu&as400cminpar@ of the White Mountains &dl 1968). Mean annual air temperatures range from less than 4" C in northern New England to 10" C in parts of southeastern New England, northern New Jersey, and northeastern Pennsylvania (Cunningham and Ciokosz 1984). Average daily minimum temperatures in January are below freezing throughout the glaciated Northeast, ranging from -18" C in northern New England to -3" C along the Atlantic coast (Lull 1968). Average daily maximum temperatures in July range from 21" to 30" C. The length ofthe bze-free period varies from less than 90 days in parts of the White Mountah, Gren Mountains, and Adirondacksto 180-210 days in coastal areas of wuthernNew England (Lull 1968). Table 1.2 summks climatic Table 1.2. Climatic data for the northeastern United States, by physiographic region (from Lull 1968). Region Mean annual Mean annual Mean freezeprecipitation snowfall _M+daily_ +sArnp2-CQ free period (cm> (cm) Jan. min. July max. (days) New England Upland 107 188 -13 27 128 New England Seaboard Lowland 109 145 -9 27 157 White Mountains 102 257 - 16 26 112 Green Mountains 107 188 -12 27 111 Adirondacks 107 272 - 14 27 114 Great Lakesa 84 190 -10 28 14-8 Glaciated Allegheny Plateau 102 163 -9 28 127 Ridge and ~dle$ 102 84 -6 29 159 Fiedmontb 112 66 -4 31 172 Coastal Plainb 114 46 -3 29 192 - - - -.-- -- -- - -- -- - -- --- - --- -- aIncludes ~lunatic data from thr St Lawrence 'balley region described in this report -- -- '~ncludes data from unglac~ated and Mew Jersey states (West Virgin~n, Maryland, and 1)elaaare) and fro1-r) uxrglaciatcd ponlons of Pennsyl-vanla --

C-J Spruce-Fa Beech-Birch-Maple Mite Ptne-Hemlock-Hardwood in] Oak-Yellow Poplar Pitch Pine-Hardwood Fig. 1.3. Major forest regj om of the glaciated Northeast (after Lull 1968 and Little 1979). Limit of Wisconsin T data for each physiographic region in the Northeast. Major Forest Regions The forests of the glaciated Northeast can be divided into five major regions (Fig. 1.3), which are differentiated according to the forest associations that dominate the upland landscape: spruce-fir, beechbirch-maple, white pine-hemlock-hardwood, oakyellow-poplar, and pitch pine-hardwood. As comparison of Figs. 1.2 and 1.3 suggests, the confition of the various forest regions is determined largely by physiography and related climatic fa&. Table 1.3 identifies the most common tree species found on upland and wetland sites in the five forest regions. Red maple swamps occur throughout the North&, but their relative abundance and floristic mmposition vary with physiography and forest region Cenerallx Cdese wetlands are most abundant in the white pine-hemlock-hardwd region and least abundant in the spmm-fir region. Eeologg~ and Distribution of Red Maple Red maple is an extremely broadly adapted species that OGCWS in both wetland and upland habitats throughout the eastern United States @owells 1965). It is found virtually everywhere east of the 100th meridian where precipitation is adequate to support tree growth (Fig. 1.4). It occurs on dry, moist, and wet soils derived from a wide variety of bedrock types, ranging from acidic granites and gneisses to basic sedimentary rocks such as limestone. It grows on dry mountain ridges, in seasonally flooded depressions with organic or mineral soils, in mesic hardwood forests, in bored conifer forests, and in southern bottomlands. Both northern and southern wetland studies characterize red maple as a moderately flood-tolerant tree (Hall and Smith 1955; Teskey and Hinckley 1978a, 197810; McKnight et al. 1981; Theriot 1988) that is most common on sites that are intermediate in wetness between permanent flooding and temporary or intermittent flooding (Buell and Wistendahl 1955; Satterlund 1960; Monk 1966; Sollers 1973; Dabel and Day 1977; Conner and Day 1982; Huenneke 1982). In the glaciated Northeast, red maple predominates in swamps where soils are saturated or flooded from late fall through early summer in most years. The Society of American Foresters (SAF) currently recognizes 90 forest cover types in the eastern United States (Eyre 1980). Redmaple is a major component (i.e., composes at least 2% of total stand basal area) in five of these types and is listed as an associated species in 63 others. It is a major or associated species in 41 of the 43 forest cover

C-J Spruce-Fa<br />

Beech-Birch-<strong>Maple</strong><br />

Mite Ptne-Hemlock-Hardwood<br />

<strong>in</strong>] Oak-Yellow Poplar<br />

Pitch P<strong>in</strong>e-Hardwood<br />

Fig. 1.3. Major forest regj om <strong>of</strong> <strong>the</strong> glaciated<br />

Nor<strong>the</strong>ast (after Lull 1968 and<br />

Little 1979).<br />

Limit <strong>of</strong> Wiscons<strong>in</strong><br />

T<br />

data for each physiographic region <strong>in</strong> <strong>the</strong> Nor<strong>the</strong>ast.<br />

Major Forest Regions<br />

The forests <strong>of</strong> <strong>the</strong> glaciated Nor<strong>the</strong>ast can be divided<br />

<strong>in</strong>to five major regions (Fig. 1.3), which are<br />

differentiated accord<strong>in</strong>g to <strong>the</strong> forest associations that<br />

dom<strong>in</strong>ate <strong>the</strong> upland landscape: spruce-fir, beechbirch-maple,<br />

white p<strong>in</strong>e-hemlock-hardwood, oakyellow-poplar,<br />

and pitch p<strong>in</strong>e-hardwood. As comparison<br />

<strong>of</strong> Figs. 1.2 and 1.3 suggests, <strong>the</strong> confition <strong>of</strong><br />

<strong>the</strong> various forest regions is determ<strong>in</strong>ed largely by<br />

physiography and related climatic fa&.<br />

Table 1.3 identifies <strong>the</strong> most common tree species<br />

found on upland and wetland sites <strong>in</strong> <strong>the</strong> five forest<br />

regions. <strong>Red</strong> maple swamps occur throughout <strong>the</strong><br />

North&, but <strong>the</strong>ir relative abundance and floristic<br />

mmposition vary with physiography and forest region<br />

Cenerallx Cdese wetlands are most abundant <strong>in</strong> <strong>the</strong><br />

white p<strong>in</strong>e-hemlock-hardwd region and least abundant<br />

<strong>in</strong> <strong>the</strong> spmm-fir region.<br />

Eeologg~ and Distribution <strong>of</strong><br />

<strong>Red</strong> <strong>Maple</strong><br />

<strong>Red</strong> maple is an extremely broadly adapted species<br />

that OGCWS <strong>in</strong> both wetland and upland habitats<br />

throughout <strong>the</strong> eastern United States @owells<br />

1965). It is found virtually everywhere east <strong>of</strong> <strong>the</strong><br />

100th meridian where precipitation is adequate to<br />

support tree growth (Fig. 1.4). It occurs on dry,<br />

moist, and wet soils derived from a wide variety <strong>of</strong><br />

bedrock types, rang<strong>in</strong>g from acidic granites and<br />

gneisses to basic sedimentary rocks such as limestone.<br />

It grows on dry mounta<strong>in</strong> ridges, <strong>in</strong> seasonally<br />

flooded depressions with organic or m<strong>in</strong>eral<br />

soils, <strong>in</strong> mesic hardwood forests, <strong>in</strong> bored conifer<br />

forests, and <strong>in</strong> sou<strong>the</strong>rn bottomlands. Both nor<strong>the</strong>rn<br />

and sou<strong>the</strong>rn wetland studies characterize red<br />

maple as a moderately flood-tolerant tree (Hall and<br />

Smith 1955; Teskey and H<strong>in</strong>ckley 1978a, 197810;<br />

McKnight et al. 1981; Theriot 1988) that is most<br />

common on sites that are <strong>in</strong>termediate <strong>in</strong> wetness<br />

between permanent flood<strong>in</strong>g and temporary or <strong>in</strong>termittent<br />

flood<strong>in</strong>g (Buell and Wistendahl 1955;<br />

Satterlund 1960; Monk 1966; Sollers 1973; Dabel<br />

and Day 1977; Conner and Day 1982; Huenneke<br />

1982). In <strong>the</strong> glaciated Nor<strong>the</strong>ast, red maple predom<strong>in</strong>ates<br />

<strong>in</strong> swamps where soils are saturated or<br />

flooded from late fall through early summer <strong>in</strong> most<br />

years.<br />

The Society <strong>of</strong> American Foresters (SAF) currently<br />

recognizes 90 forest cover types <strong>in</strong> <strong>the</strong> eastern<br />

United States (Eyre 1980). <strong>Red</strong>maple is a major<br />

component (i.e., composes at least 2% <strong>of</strong> total<br />

stand basal area) <strong>in</strong> five <strong>of</strong> <strong>the</strong>se types and is listed<br />

as an associated species <strong>in</strong> 63 o<strong>the</strong>rs. It is a major<br />

or associated species <strong>in</strong> 41 <strong>of</strong> <strong>the</strong> 43 forest cover

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