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Behavioural heterogeneity<br />

and human cooperation<br />

Maintaining large-scale cooperation requires incentives to<br />

overcome cooperation dilemmas in which freeriders enjoy<br />

public goods created by others without bearing the costs.<br />

Literature suggests (i) that conditional cooperation and<br />

costly punishment can stabilise cooperation, and (ii) that<br />

group selection based on culture provides larger scope for<br />

the evolution of these traits. However, most evidence on the<br />

former remains a major empirical challenge when applied<br />

in the real world. Evaluation of the latter could be improved<br />

by measuring culture through behavioural experiments and<br />

by estimating cultural differentiation among neighbouring<br />

groups of small-scale societies. In this study, we employed<br />

a variety of behavioural experiments, household questionnaires,<br />

and community surveys to measure these behaviours<br />

as well as other factors, and then study their effect<br />

on the outcomes of forest commons management. The<br />

study area covered 49 groups of the Bale Oromo people in<br />

Ethiopia.<br />

Our results show that high shares of conditional cooperators<br />

in a group have a significantly positive effect on the<br />

outcomes, even after controlling for conventional factors.<br />

Costly monitoring is a key mechanism by which conditional<br />

cooperators sustain cooperation. Our preliminary investigations<br />

further show that freely conferred deference on the<br />

group leader partially accounts for group level differences in<br />

conditional cooperation. The investigations also show that<br />

cultural group selection, as estimated through cultural differentiation<br />

among 77 pairs of neighbouring small-scale<br />

societies, provides scope for the evolution of costly cooperative<br />

traits. A detailed examination of the punishment<br />

behaviour of a group leader in response to deviations from<br />

co-operation norms reveals substantial variations. Normdriven<br />

leaders who explicitly punish deviations from the<br />

conditional cooperation norm have a positive but insignificant<br />

effect on the outcomes. In contrast, spiteful leaders<br />

who punish cooperators have a significantly negative effect.<br />

Furthermore, we investigated how behavioural heterogeneity<br />

and beliefs interact in voluntary cooperation. We<br />

find that most Bale Oromo conditional cooperators exhibit<br />

‘‘altruistic’’ bias, contributing slightly more than the partner<br />

player does. This challenges the hypothesis that imperfect<br />

conditional cooperation causes decline in cooperation over<br />

time. Our results reveal that behavioural heterogeneity is<br />

a highly significant predictor of voluntary cooperation and<br />

that beliefs have a positive effect on cooperation only when<br />

a player is a conditional cooperator.<br />

76<br />

Research fellow<br />

Devesh Rustagi, <strong>ETH</strong> Zurich, Switzerland<br />

Supervisors<br />

Stefanie Engel, <strong>ETH</strong> Zurich, Switzerland;<br />

Michael Kosfeld, University of Frankfurt, Germany<br />

Capacity development<br />

Research fellowships<br />

Collaborators<br />

Bruce Cambell, CIFOR, Indonesia;<br />

Franz Gatzweiler and Günther Manske,<br />

ZEF, University of Bonn, Germany;<br />

Martin Neumann, GTZ Sustainable Utilization of Natural<br />

Resources for Food Security (SUN), Oromia, Ethiopia;<br />

Gurara Gebissa, Regional State of Oromia Forest<br />

Enterprises Supervising Agency, Ethiopia<br />

Duration<br />

September 2008 – December 2009<br />

Economic experiments used to analyse the prevalence<br />

of social preferences in Ethiopian communities

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