Gymnocystal development
Gymnocystal development
Gymnocystal development
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Research and Status<br />
of Taiwan Species<br />
Biodiversity<br />
Taichung<br />
15-16 August 2008
Bryozoans<br />
range from<br />
the intertidal<br />
to the deep<br />
sea and can<br />
form<br />
significant<br />
components<br />
of rock-wall<br />
assemblages.
Mostly bryozoan hash
Washed<br />
bryomol<br />
gravel
Some bryozoans are habitat-formers —<br />
like species of Celleporaria ...
But what about<br />
Taiwan?
Liu, Yin & Ma<br />
“Biology of Marine-<br />
Fouling Bryozoans in<br />
the Coastal Waters of<br />
China”<br />
Beijing 2001<br />
Lists all fouling coastal<br />
bryozoans of western<br />
Taiwan Strait
Hard and soft shores<br />
along the Strait of<br />
Taiwan on the west coast
Hard and soft shores along<br />
the Pacific east coast
Celleporaria pilaefera<br />
Hippoporina indica<br />
Watersipora subtorquata<br />
Schizoporella errata
Keelung<br />
Total<br />
expected<br />
bryofauna<br />
>600<br />
species<br />
EEZ<br />
boundary<br />
Green<br />
Island
Collection of beach-cast coral<br />
rubble made in October 2005<br />
Keelung<br />
City
Cryptofaunal bryozoans in coral<br />
environments<br />
Within reefs:<br />
hidden encrusters<br />
cavity dwellers<br />
cavity fillers<br />
dead-coral veneerers<br />
Between reefs:<br />
sediment binders and trappers<br />
sediment stabilisers<br />
Hence, bryozoans can contribute to reef build-up, especially those<br />
species that engage in self-overgrowth or which grow over other<br />
organisms. Such bryozoans tend to become a permanent part of the<br />
calcareous mass that comprises a reef, acting as accessory framebuilders.
Class Gymnolaemata<br />
Order Cheilostomata<br />
Order Ctenostomata
Class Gymnolaemata<br />
Class Phylactolaemata<br />
Order Cheilostomata<br />
Order Plumatellida<br />
Order Ctenostomata
Class Gymnolaemata<br />
Class Phylactolaemata<br />
Order Cheilostomata<br />
Order Plumatellida<br />
Class Stenolaemata<br />
Order Ctenostomata<br />
Order Cyclostomata
Class Gymnolaemata<br />
Class Stenolaemata<br />
Order Cheilostomata<br />
Order Ctenostomata<br />
Order Cyclostomata
Suborder Inovicellina<br />
Superfamily Aeteoidea<br />
Family Aeteidae<br />
AETEIDAE<br />
Aetea sp.
gymnocyst<br />
membranous<br />
dddd<br />
frontal<br />
dddd<br />
wall<br />
d<br />
<strong>Gymnocystal</strong> <strong>development</strong><br />
Cryptocystal <strong>development</strong><br />
In anascan<br />
cheilostomes<br />
cryptocyst<br />
hypostegal<br />
coelom
gymnocyst<br />
membranous<br />
dddd<br />
frontal<br />
dddd<br />
wall<br />
d<br />
<strong>Gymnocystal</strong> <strong>development</strong><br />
in anascan cheilostomes<br />
Cryptocystal <strong>development</strong><br />
cryptocyst<br />
hypostegal<br />
coelom
ANTROPORIDAE<br />
Antropora sp.<br />
Antropora minor<br />
(Hincks, 1880)
Ascophorine frontal shields<br />
Spinocystal shield<br />
<strong>Gymnocystal</strong> shield
CRIBRILINIDAE<br />
CRIBRILINIDAE<br />
Puellina egretta Hayward & Ryland, 1995<br />
Puellina vulgaris Ryland & Hayward, 1992
HIPPOTHOIDAE<br />
Hippothoa calciophilia Gordon, 1984
TRYPOSTEGIDAE<br />
Trypostega johnsoulei Tilbrook, 2006
Ascophorine frontal shields<br />
Umbonuloid shield<br />
Lepralioid shield
ARACHNOPUSIIDAE<br />
Poricella spathulata (Canu & Bassler, 1929)
LEPRALIELLIDAE<br />
Celleporaria sp.
LEPRALIELLIDAE<br />
Drepanophora gutta<br />
Tilbrook, Hayward & Gordon 2006
ROMANCHEINIDAE<br />
Escharoides longirostris Dumont, 1981
BITECTIPORIDAE<br />
Metroperiella sp.
SMITTINIDAE<br />
Pleurocodonellina<br />
signata (Waters, 1889)<br />
Pleurocodonellina soulesi<br />
Scholz & Cusi, 1991
WATERSIPORIDAE<br />
Watersipora edmondsoni Soule & Soule, 1968
SCHIZOPORELLIDAE<br />
Hippomenella cf. lateralis (MacGillivray, 1891)
ESCHARINIDAE<br />
Pesanserina sp.
GIGANTOPORIDAE<br />
Cosciniopsis lonchaea (Busk, 1884)
MICROPORELLIDAE<br />
Calloporina sigillata Canu & Bassler, 1929
MICROPORELLIDAE<br />
Microporella orientalis Harmer, 1957
CREPIDACANTHIDAE<br />
Crepidacantha grandis Canu & Bassler, 1929
MICROPORELLIDAE<br />
Fenestrulina sp.
CELLEPORIDAE<br />
Predanophora n. sp.
PHIDOLOPORIDAE<br />
Lifuella porelliformis<br />
(Kirkpatrick, 1888)<br />
Lifuella lepralielloidum<br />
Liu in Liu, Yin & Ma, 2001
PHIDOLOPORIDAE<br />
Schedocleidochasma porcellaniforme<br />
Soule, Soule & Chaney, 1991
PHIDOLOPORIDAE<br />
Rhynchozoon sp.
ANNECTOCYMIDAE<br />
Annectocyma sp.
LICHENOPORIDAE<br />
Disporella sp.
FINAL TALLY OF<br />
TAIWANESE BRYOZOA?<br />
Green Island<br />
: 31 species<br />
Present study : 44 species<br />
Combined list, corrected for overlap) : 72 species<br />
Estimate for Taiwanese EEZ : >600 species