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Strychnos 1990 - 2004 - Crops for the Future

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Author<br />

Title<br />

Year<br />

Source title<br />

Reference<br />

Vanveenendaal, W. L. H. and R. W. Denouter<br />

Development of included phloem and organization of <strong>the</strong> phloem network in <strong>the</strong> st<br />

1993<br />

Iawa Journal<br />

14(3): 253-265<br />

Abstract<br />

The development of <strong>the</strong> diffuse included phloem strands in <strong>Strychnos</strong> millepunctata Leeuwenberg is<br />

described and compared with that in some o<strong>the</strong>r <strong>Strychnos</strong> species. - The phloem strands are<br />

produced outwards by small segments of <strong>the</strong> vascular cambium which function differently. In <strong>the</strong>se<br />

segments <strong>the</strong> secondary xylem production inwards is retarted and gradually stops, <strong>the</strong> secondary<br />

phloem production outwards increases. Thus <strong>the</strong> vascular cambium cylinder <strong>for</strong>ms depressions.<br />

These cambium segments become detached from <strong>the</strong> rest of <strong>the</strong> vascular cambium. The future strand<br />

sinks into <strong>the</strong> secondary xylem and is bridged over by a complementary cambium which is <strong>for</strong>med in<br />

<strong>the</strong> outer border of <strong>the</strong> strand and joins <strong>the</strong> vascular cambial cylinder. The complementary cambium<br />

now starts with <strong>the</strong> production of secondary xylem inwards and secondary phloem outwards like <strong>the</strong><br />

rest of <strong>the</strong> vascular cambium in die once more closed cambial cylinder. - The composition of <strong>the</strong><br />

secondary phloem in <strong>the</strong> strands is different from both <strong>the</strong> secondary and primary external phloem<br />

(phloem outside <strong>the</strong> cylinder of vascular cambium). Radial rows of cells are present within <strong>the</strong><br />

included phloem strands which continue into <strong>the</strong> later <strong>for</strong>med secondary xylem. One- to five-seriate<br />

rays traverse <strong>the</strong> strands. The embedded cambial segment near <strong>the</strong> inner border of <strong>the</strong> strand<br />

remains active. The crushing of <strong>the</strong> phloem takes place near <strong>the</strong> outer border Of <strong>the</strong> strand, <strong>for</strong>ming<br />

cap-like tissues of disorganised cells. Internal phloem is regularly renewed by cambial activity. The<br />

internal cambium produces secondary phloem inwards. The composition of this phloem resembles<br />

that of <strong>the</strong> included phloem strands. - Inter-connections between included phloem strands, as well as<br />

anastomoses of <strong>the</strong>se strands with both secondary external and internal phloem, has been<br />

demonstrated. Toge<strong>the</strong>r <strong>the</strong>se three phloem types <strong>for</strong>m a three-dimensional secondary phloem<br />

n e t w o r k .<br />

Author<br />

Title<br />

Year<br />

Source title<br />

Reference<br />

Abstract<br />

Amat, M. and J. Bosch<br />

Studies on <strong>the</strong> syn<strong>the</strong>sis of pentacyclic <strong>Strychnos</strong> indole alkaloids - closure of e-ri<br />

1992<br />

Journal of Organic Chemistry<br />

57(21): 5792-5796<br />

-

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