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On the Ecology of Mountainous Forests in a Changing Climate: A ...

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Behaviour <strong>of</strong> FORCLIM along a transect <strong>in</strong> <strong>the</strong> European Alps 99<br />

4.2 FORCLIM-S<br />

The buildup <strong>of</strong> soil organic matter was evaluated at six sites along a gradient <strong>of</strong> <strong>in</strong>creas<strong>in</strong>g<br />

actual evapotranspiration (Bever, north-fac<strong>in</strong>g slope; Davos; Sion; Bever, south-fac<strong>in</strong>g<br />

slope; Bern; Locarno; cf. Tab. 4.1). It was assumed that <strong>the</strong>re is no organic material at<br />

<strong>the</strong> beg<strong>in</strong>n<strong>in</strong>g <strong>of</strong> <strong>the</strong> simulations and that <strong>the</strong>re is a constant annual <strong>in</strong>put <strong>of</strong> litter <strong>in</strong>to <strong>the</strong><br />

system (Tab. 4.4). The simulations were run until <strong>the</strong> steady state <strong>of</strong> soil organic matter<br />

was reached.<br />

Bever (north fac<strong>in</strong>g slope), ForClim-S<br />

400<br />

Organic matter [t/ha]<br />

Available nitrogen [kg/ha]<br />

300<br />

200<br />

100<br />

0<br />

0<br />

50<br />

100<br />

150<br />

200<br />

250<br />

300<br />

uAvN<br />

LOM<br />

HOM<br />

SOM<br />

Simulation time [years]<br />

Fig. 4.5: Buildup <strong>of</strong> soil organic matter content on a north-fac<strong>in</strong>g slope at <strong>the</strong> site Bever<br />

as simulated by FORCLIM-S. LOM: litter organic matter; HOM: humus organic matter;<br />

SOM: total soil organic matter, SOM = LOM + HOM; uAvN: nitrogen available for plant<br />

growth. Litter <strong>in</strong>put is from Tab. 4.4, and actual evapotranspiration is from Tab. 4.1.<br />

Fig. 4.5 shows typical simulation results obta<strong>in</strong>ed from FORCLIM-S for a nor<strong>the</strong>rn slope<br />

at <strong>the</strong> site Bever. The steady state <strong>of</strong> belowground organic matter is reached with<strong>in</strong> 200-<br />

250 years, and Tab. 4.3 gives an overview <strong>of</strong> <strong>the</strong> steady-state results obta<strong>in</strong>ed at <strong>the</strong> six<br />

sites.<br />

As mentioned <strong>in</strong> <strong>the</strong> section on <strong>the</strong> assumptions <strong>of</strong> FORCLIM-S, both LINKAGES and<br />

FORCLIM-S lack a carbon pool with a turnover rate <strong>in</strong> <strong>the</strong> order <strong>of</strong> 1000 years (Parton et<br />

al. 1987, Verberne et al. 1990). Hence <strong>the</strong> steady state <strong>of</strong> belowground organic matter is<br />

reached too fast and is slightly too low <strong>in</strong> <strong>the</strong>se models (cf. Fig. 4.5). A hypo<strong>the</strong>sized<br />

“slow” compartment would be small and would react much more slowly to climatic<br />

change than <strong>the</strong> species composition. Moreover, with<strong>in</strong> <strong>the</strong> next few hundred years <strong>the</strong>

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