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On the Ecology of Mountainous Forests in a Changing Climate: A ...

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The forest model FORCLIM 77<br />

NITROGEN AVAILABILITY FOR PLANT GROWTH<br />

The amount <strong>of</strong> nitrogen available for plant growth (uAvN, Eq. 3.62) is calculated as an<br />

output variable <strong>of</strong> <strong>the</strong> FORCLIM-S model from <strong>the</strong> net nitrogen immobilization <strong>of</strong> all <strong>the</strong><br />

nLC litter cohorts (Eq. 3.61) and <strong>the</strong> nitrogen m<strong>in</strong>eralization rate (Eq. 3.57). uAvN is<br />

not a state variable because it is assumed that <strong>the</strong> available nitrogen not used by <strong>the</strong> plants<br />

<strong>in</strong> a given year leaves <strong>the</strong> system ei<strong>the</strong>r by streamflow or as volatile nitrogen compounds.<br />

Eq. 3.62 also <strong>in</strong>cludes <strong>the</strong> atmospheric deposition <strong>of</strong> soluble N compounds (kNAtm).<br />

gLImmob =<br />

nLC<br />

∑<br />

c = 1<br />

gNetImmob c<br />

(3.61)<br />

uAvN = MAX ∆HN<br />

∆t<br />

– gLImmob, 0 + kNAtm (3.62)<br />

3.3.3 FORCLIM-E: A model <strong>of</strong> <strong>the</strong> abiotic environment<br />

All <strong>the</strong> equations used <strong>in</strong> <strong>the</strong> submodel FORCLIM-E were described previously <strong>in</strong> our<br />

analysis <strong>of</strong> <strong>the</strong> sensitivity <strong>of</strong> forest gap models to climate parametrization schemes<br />

(Fischl<strong>in</strong> et al. 1994). Thus, only <strong>the</strong> modifications made to <strong>the</strong>se equations are presented<br />

and discussed here, us<strong>in</strong>g <strong>the</strong> same notational conventions as <strong>in</strong> Fischl<strong>in</strong> et al. (1994).<br />

GENERATION OF WEATHER DATA<br />

Cross-correlated variates <strong>of</strong> monthly mean temperature (T) and monthly precipitation sum<br />

(P) are generated us<strong>in</strong>g <strong>the</strong> follow<strong>in</strong>g method: First, we note that <strong>the</strong> long-term means (µ)<br />

and standard deviations (σ) <strong>of</strong> <strong>the</strong>se variables may be written as vectors (Eq. 3.63), and<br />

<strong>the</strong>ir cross-correlations (r) as a matrix (Eq. 3.64):<br />

µ m,l =<br />

µ T,m,l<br />

µ P,m,l<br />

σ m,l =<br />

σ T,m,l<br />

σ P,m,l<br />

(3.63)<br />

R m,l =<br />

r TT,m,l<br />

r TP,m,l<br />

r PT,m,l r PP,m,l<br />

=<br />

1 r TP,m,l<br />

r PT,m,l 1<br />

(3.64)<br />

where <strong>the</strong> subscript m stands for a given month, and l for a location. The covariance<br />

matrix <strong>of</strong> <strong>the</strong> two variables is given <strong>in</strong> Eq. 3.65 (Flury & Riedwyl 1983):

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