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On the Ecology of Mountainous Forests in a Changing Climate: A ...

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158 Chapter 6<br />

F<strong>in</strong>ally, most climate scenarios give us an <strong>in</strong>dication about what <strong>the</strong> change <strong>in</strong> a given<br />

wea<strong>the</strong>r variable will be by a certa<strong>in</strong> po<strong>in</strong>t <strong>in</strong> <strong>the</strong> future. However, <strong>in</strong> most scenarios it is<br />

not known how this change will be realized. Most impact studies so far have assumed<br />

that climate changes l<strong>in</strong>early (Solomon 1986, Pastor & Post 1988, Kienast 1991); o<strong>the</strong>r<br />

authors have studied <strong>the</strong> response <strong>of</strong> forest models to an <strong>in</strong>stantaneous climatic change,<br />

i.e. its step response (Fischl<strong>in</strong> et al. 1994, Bugmann & Fischl<strong>in</strong> 1994). Thus, <strong>the</strong>re arises<br />

<strong>the</strong> question how sensitive <strong>the</strong> behaviour <strong>of</strong> forest gap models is to <strong>the</strong> assumptions on<br />

<strong>the</strong> nature <strong>of</strong> transient climatic change.<br />

6.2.1 Material & methods<br />

Forest models<br />

Five forest gap models are used <strong>in</strong> this study: The first model, FORECE (Kienast 1987),<br />

is a conventional gap model that was analyzed <strong>in</strong> detail <strong>in</strong> chapter 2. The second model,<br />

FORCLIM 1.1 (Bugmann & Fischl<strong>in</strong> 1994), is a simplified descendant <strong>of</strong> FORECE and<br />

comprises only <strong>the</strong> most fundamental ecological processes (cf. section 2.3.1); <strong>the</strong> formulation<br />

<strong>of</strong> climatic factors <strong>in</strong> FORCLIM 1.1 is done <strong>in</strong> <strong>the</strong> same way as <strong>in</strong> FORECE. The<br />

third model, FORCLIM 1.3 (Bugmann & Fischl<strong>in</strong> 1994), was developed from FORCLIM<br />

1.1 by alter<strong>in</strong>g <strong>the</strong> ma<strong>the</strong>matical formulations <strong>of</strong> <strong>the</strong> climatic factors (cf. section 2.3.2 and<br />

Fischl<strong>in</strong> et al. 1994). The fourth model, FORCLIM-E/P, is one variant <strong>of</strong> <strong>the</strong> model developed<br />

<strong>in</strong> chapter 3; it differs from FORCLIM 1.3 concern<strong>in</strong>g <strong>the</strong> formulation <strong>of</strong> ecological<br />

factors and <strong>of</strong> climatic <strong>in</strong>fluences. The last model is FORCLIM-E/P/S, which <strong>in</strong>corporates<br />

also belowground carbon and nitrogen turnover (chapter 3).<br />

Study sites and steady state climate scenarios<br />

Six sites were selected along a climatological and at <strong>the</strong> same time altitud<strong>in</strong>al gradient,<br />

rang<strong>in</strong>g from above <strong>the</strong> current alp<strong>in</strong>e timberl<strong>in</strong>e to central alp<strong>in</strong>e valleys close to <strong>the</strong> dry<br />

timberl<strong>in</strong>e. In a previous study <strong>of</strong> possible impacts <strong>of</strong> climatic change on forests, Kienast<br />

(1991) presented simulation results from <strong>the</strong> sites St. Gotthard, Airolo, and Sion along<br />

with results from 15 o<strong>the</strong>r sites. To allow for a comparison <strong>of</strong> <strong>the</strong> results from <strong>the</strong><br />

Kienast (1991) study with <strong>the</strong> ones obta<strong>in</strong>ed here, <strong>the</strong>se three sites will be used as well.<br />

Gyalistras et al. (1994) developed a methodology to relate large-scale temperature and<br />

pressure anomalies to local wea<strong>the</strong>r anomalies by means <strong>of</strong> pr<strong>in</strong>cipal component analysis<br />

and canonical correlation analysis (“Downscal<strong>in</strong>g”). Based on <strong>the</strong> results obta<strong>in</strong>ed by this<br />

technique for <strong>the</strong> sites Bever, Davos, and Bern, Bugmann & Fischl<strong>in</strong> (1994) performed

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