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On the Ecology of Mountainous Forests in a Changing Climate: A ...

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136 Chapter 5<br />

FORECE<br />

Fagus silvatica<br />

T (°C)<br />

P (mm/yr)<br />

FORC LIM-E/P<br />

FORC LIM-E/P/S<br />

T (°C)<br />

T (°C)<br />

P (mm/yr)<br />

P (mm/yr)<br />

Fig. 5.6: Contour plot <strong>of</strong> Fagus silvatica biomass [t/ha] as simulated by FORECE (top),<br />

FORCLIM-E/P (bottom left), and FORCLIM-E/P/S (bottom right).<br />

Similar steep gradients as observed with P. excelsa and P. cembra occur with F. silvatica<br />

<strong>in</strong> <strong>the</strong> FORECE model (Fig. 5.6): There is an abrupt decl<strong>in</strong>e at T l ≈ 5 °C, which is<br />

located <strong>in</strong> <strong>the</strong> middle <strong>of</strong> <strong>the</strong> upper montane beech-silver fir zone (Fig. 5.1). Aga<strong>in</strong>, both<br />

variants <strong>of</strong> FORCLIM produce smoo<strong>the</strong>r and more realistic gradients; specifically, <strong>the</strong><br />

species extends down to T l ≈ 4 °C, where <strong>the</strong> transition between montane and subalp<strong>in</strong>e<br />

forests should occur (Fig. 5.1). Moreover, <strong>in</strong> FORCLIM F. silvatica extends down to a<br />

precipitation <strong>of</strong> about 800 mm, which is plausible as well (Fig. 5.6).<br />

Nei<strong>the</strong>r FORECE nor FORCLIM are capable <strong>of</strong> simulat<strong>in</strong>g <strong>the</strong> transition from beech forests<br />

to <strong>in</strong>subrian and mediterranean forests dom<strong>in</strong>ated by oak (Quercus spp.) and chestnut (C.<br />

sativa), which should occur at T l ≈ 10 °C (Fig. 5.6). In both models, Fagus silvatica

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