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On the Ecology of Mountainous Forests in a Changing Climate: A ...

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Parameter sensitivity & model validation 133<br />

Total biomass (FORECE)<br />

t/ha<br />

500<br />

400<br />

300<br />

200<br />

100<br />

12<br />

10<br />

8<br />

6<br />

T (°C)<br />

4<br />

2<br />

0<br />

40<br />

90 140 190 0<br />

P (cm/yr)<br />

Total biomass (E/P)<br />

Total biomass (E/P/S)<br />

t/ha<br />

t/ha<br />

500<br />

600<br />

400<br />

500<br />

300<br />

200<br />

100<br />

400<br />

300<br />

200<br />

100<br />

12<br />

10<br />

8<br />

6<br />

T (°C)<br />

4<br />

2<br />

0<br />

40<br />

0<br />

90 140 190 P (cm/yr)<br />

12<br />

10<br />

8<br />

6<br />

4<br />

T (°C)<br />

2<br />

0<br />

40<br />

0<br />

90 140 190 P (cm/yr)<br />

Fig. 5.3: Total aboveground biomass as simulated by <strong>the</strong> three models FORECE, FOR-<br />

CLIM-E/P and FORCLIM-E/P/S <strong>in</strong> a space spanned by <strong>the</strong> annual precipitation sum (P) and<br />

<strong>the</strong> annual mean temperature (T).<br />

istic. However, it should be noted that nitrogen availability has been <strong>in</strong>troduced <strong>in</strong> FOR-<br />

CLIM as a substitute <strong>of</strong> SOILQ, and that <strong>the</strong> assumption <strong>of</strong> a constant supply <strong>of</strong> nitrogen<br />

(100 kg/ha) <strong>in</strong> FORCLIM-E/P is equally unrealistic; yet this model does not produce <strong>the</strong><br />

anomaly evident from FORECE (Fig 5.3).<br />

The major difference <strong>of</strong> aboveground biomass between <strong>the</strong> two FORCLIM variants is that<br />

<strong>in</strong> FORCLIM-E/P/S nitrogen availability rises above 100 kg/ha at higher temperatures,<br />

and total biomass <strong>in</strong>creases above <strong>the</strong> level reached by FORCLIM-E/P (Fig. 5.3, cf.<br />

Fig. 5.8).

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