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On the Ecology of Mountainous Forests in a Changing Climate: A ...

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Behaviour <strong>of</strong> FORCLIM along a transect <strong>in</strong> <strong>the</strong> European Alps 117<br />

this would not be true for much larger ∆t where autocorrelation becomes negligible. The<br />

choice <strong>of</strong> n = 200 and ∆t = 150 seems to provide a good compromise between <strong>the</strong> simulation<br />

time needed and <strong>the</strong> accuracy <strong>of</strong> <strong>the</strong> estimation; moreover, <strong>the</strong>se data conform to <strong>the</strong><br />

considerations <strong>in</strong> <strong>the</strong> sections 2.2.1 (autocorrelation) and 2.2.2 (sample size).<br />

SIMILARITY OF INDEPENDENT ESTIMATES OF THE SAME STEADY STATE<br />

The histograms <strong>of</strong> <strong>the</strong> distribution <strong>of</strong> <strong>the</strong> 200 PS coefficients obta<strong>in</strong>ed from 400 simulation<br />

runs at <strong>the</strong> three sites conform to <strong>the</strong> hypo<strong>the</strong>sis formulated above (Fig. 4.15,<br />

Tab. 4.6): In <strong>the</strong> species-poor Larici-Piceetum at Davos (Ellenberg & Klötzli 1972), <strong>the</strong><br />

PS coefficients are considerably higher than <strong>in</strong> <strong>the</strong> diverse forests <strong>of</strong> Airolo and Bern.<br />

The reason for <strong>the</strong> decreas<strong>in</strong>g PS coefficients with <strong>in</strong>creas<strong>in</strong>g species diversity can be expla<strong>in</strong>ed<br />

by consider<strong>in</strong>g <strong>the</strong> averages <strong>of</strong> <strong>the</strong> estimated species-specific biomasses and <strong>the</strong>ir<br />

coefficients <strong>of</strong> variation (CV, Zar 1984) from <strong>the</strong> 400 steady states estimated at each site.<br />

Frequency<br />

Bern<br />

40<br />

35<br />

30<br />

25<br />

20<br />

15<br />

10<br />

5<br />

0<br />

.8 .825 .85 .875 .9 .925 .95 .975 1<br />

PS<br />

Airolo<br />

40<br />

35<br />

30<br />

25<br />

20<br />

15<br />

10<br />

5<br />

0<br />

.8 .825 .85 .875 .9 .925 .95 .975 1<br />

PS<br />

Frequency<br />

Davos<br />

40<br />

35<br />

30<br />

25<br />

20<br />

15<br />

10<br />

5<br />

0<br />

.8 .825 .85 .875 .9 .925 .95 .975 1<br />

PS<br />

Fig. 4.15: Frequency distribution <strong>of</strong> <strong>the</strong> percentage similarity coefficients (PS) from 200<br />

pairs <strong>of</strong> equilibrium states estimated at <strong>the</strong> sites Bern, Airolo, and Davos, us<strong>in</strong>g n=200 and<br />

∆t=150 years. The distributions at <strong>the</strong> sites Bern and Airolo are not significantly different<br />

from each o<strong>the</strong>r (p = 0.12, Kolmogorov-Smirnov test, Zar 1984).

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