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Alafia River Minimum Flows and Levels - Southwest Florida Water ...

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subsidy of water <strong>and</strong> nutrients that supports high rates of primary production in<br />

river floodplains (Conner <strong>and</strong> Day 1976, Brinson et al. 1981). This primary<br />

production produces large amounts of organic detritus, which is critical to food<br />

webs on the floodplain <strong>and</strong> within the river channel (Vannote et al. 1980, Gregory<br />

et al. 1991). Floodplain inundation also contributes to other physical-chemical<br />

processes that can affect biological production, uptake <strong>and</strong> transformation of<br />

macro-nutrients (Kuensler 1989, Walbridge <strong>and</strong> Lockaby 1994).<br />

Soils in river floodplains exhibit physical <strong>and</strong> chemical properties that are<br />

important to the overall function of the river ecosystem (Wharton et al. 1982,<br />

Stanturf <strong>and</strong> Schenholtz 1998). Anaerobic soil conditions can persist in areas<br />

where river flooding or soil saturation is of sufficient depth <strong>and</strong> duration. The<br />

decomposition of organic matter is much slower in anaerobic environments, <strong>and</strong><br />

mucky or peaty organic soils can develop in saturated or inundated floodplain<br />

zones (Tate 1980, Brown et al. 1990). Although these soils may dry out on a<br />

seasonal basis, typically long hydroperiods contribute to their high organic<br />

content. Plant species that grow on flooded, organic soils are tolerant of anoxic<br />

conditions <strong>and</strong> the physical structure of these soils (Hook <strong>and</strong> Brown 1973,<br />

McKevlin et al. 1998). Such adaptations can be an important selective<br />

mechanism that determines plant community composition. Because changes in<br />

river hydrology can potentially effect the distribution <strong>and</strong> characteristics of<br />

floodplain soils, soil distributions <strong>and</strong> their relationship to river hydrology are<br />

routinely investigated as part of MFL determinations for District rivers.<br />

Compared to instream evaluations of MFL requirements, there has been<br />

relatively little work done on river flows necessary for meeting the requirements<br />

of floodplain species, communities or functions. Our work on the upper Peace<br />

<strong>and</strong> <strong>Alafia</strong> <strong>River</strong>s suggests that direct <strong>and</strong> continuous inundation of floodplain<br />

wetl<strong>and</strong>s by river flows is in many cases not sufficient to meet the published<br />

inundation needs of the dominant species found in the wetl<strong>and</strong>s. There are<br />

probably several reasons for this apparent inconsistency. Some floodplain<br />

systems are likely to include seepage wetl<strong>and</strong>s, dependent on hydrologic<br />

processes other than direct inundation from the river. Other wetl<strong>and</strong>s may occur<br />

in depressional areas where water is retained after subsidence of river flows.<br />

The District's approach to protection of flows associated with floodplain habitats,<br />

communities <strong>and</strong> functions involves consideration of the frequency <strong>and</strong> duration<br />

of direct connection between the river channel <strong>and</strong> the floodplain. As part of this<br />

process, plant communities <strong>and</strong> soils are identified across the river floodplain at a<br />

number of sites, <strong>and</strong> periods of inundation/connection with the river are<br />

reconstructed on an annual or seasonal basis. These data are used to<br />

characterize the frequency <strong>and</strong> duration of direct connection/ inundation of these<br />

communities to or by the river <strong>and</strong> to develop criteria for minimum flow<br />

development.<br />

3-7

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