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Boyer diss 2009 1046..

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is not convoluted like many other sutures; however, the contact between these particular<br />

bones frequently looks this way in various other taxa (e.g., tenrecs). Furthermore, two<br />

features (only one could be photo-documented) of this contact strongly suggest it is a<br />

suture: (1) the form of the discontinuity between the alisphenoid and orbitosphenoid is<br />

revealed by the absence (i.e., broken condition) of the orbitosphenoid in the region of<br />

interest. Instead of appearing “crack-like” and planar, the discontinuity is dished like a<br />

sutural contact (Fig. 2.24: 102). (2) Even though the discontinuity is not convoluted like<br />

other sutures, it is still more complex than would be expected for a crack created by<br />

brittle deformation. Identification of this final suture helps interpret cranial foraminal<br />

patterns. Having recognized this suture on MNHN CR 965, it becomes apparent that it<br />

has a slightly different course than that preserved on the left side of MNHN CR 125<br />

connecting foramina 90 and 93 (Fig. 2.20D’). The latter suture would likely have<br />

resulted in the orbitosphenoid forming most of the medial wall of the sphenorbital fissure.<br />

Cranial foramina.— Foramina for the trigeminal nerve’s mandibular division<br />

(Figs. 2.22, 25: 103), for its combined maxillary and ophthalmic divisions (Figs. 2.22-25:<br />

104), and for the optic nerve (Figs. 2.22, 24-25: 105) are clearly visible and traceable to<br />

endocranial space. There are a number of small foramina representing sinus drainage<br />

from the lateral aspects of the alisphenoid and the lateral aspect of the orbitosphenoid<br />

(Fig. 2.23, 24: 106). These cannot be mistaken for cranial nerve foramina because they<br />

are not bilaterally present in some cases or do not lead to the endocranium in other cases.<br />

This is also true for the “suboptic foramen” located posteroventral to the optic foramen<br />

(Figs. 2.22, 24: 107). It appears to lead into the trabecular space of the orbitosphenoid<br />

and probably communicates directly with the blood sinus foramina on the opposite side.<br />

63

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