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Boyer diss 2009 1046..

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is roughly 2.8 mm long. Medial to the promontorium is a groove leading to a foramen<br />

that perforates the medial process of the bulla at its medial point of termination, and one<br />

that perforates the promontorium itself at its lateral point of termination. This feature has<br />

been interpreted as the vestibular aqueduct by Szalay et al. (1987) but it is clearly the<br />

tympanic canaliculus foramen and groove, as discussed for other specimens earlier<br />

(MacPhee, 1981) (Fig. 2.21C’: 94-95). This morphology is clearly present in most other<br />

plesiadapid specimens preserving the relevant anatomy, as discussed and illustrated<br />

above. The hypoglossal canal appears septate, and split into two foramina, as in<br />

Pronothodectes.<br />

Morphology of cranial bones.– Some of the most critical information relating to<br />

the structure of the basicranium in this specimen has been lost: What remained of the<br />

medial process of the left petrosal when Gingerich (1976: Pl. 8c) photographed the<br />

specimen sometime prior to 1974 is now gone. However, a cast recently made from an<br />

old mold housed in the MNHN, retains the medial process. This cast also reveals that<br />

MNHN CR 125 was broken at the junction of medial tympanic process with pars<br />

cochlearis even before the medial process was lost (demonstrating that this specimen was<br />

never substantially better preserved than the Pellouin skull anyway).<br />

The promontoria of MNHN CR 125 conform well to the description by Gingerich<br />

(1976). As noted above, however, neither the posterior carotid foramen nor the laterally<br />

positioned g1 groove for the internal carotid plexus has been photographically illustrated<br />

previously. Figure 2.21B shows this morphology. Unlike other P. tricuspidens<br />

promontoria (see below) and unlike many other plesiadapid petrosals (see above),<br />

61

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