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Boyer diss 2009 1046..

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and projects straight ventrally by 1.35 mm. It is lateral to the postglenoid foramen (Fig.<br />

2.15: 63).<br />

Petrosal.— The pars cochlearis of both petrosals are preserved, as well as some<br />

of the septa and tympanic processes that attach to them (Figs. 2.15, 16). On the right<br />

side, a bit of the pars canalicularis is visible. The posterior septum is either broken, or<br />

differently configured than in other plesiadapid specimens, such that no evidence of the<br />

g1 groove is present. Because the posterior septum actually appears fairly complete, it<br />

may be that the internal carotid plexus did not go through the middle ear, and took a<br />

different route in this taxon (see further discussion below). The lateral aspects of the<br />

promontoria are obscured so that the presence of the g2 and g5 grooves and anterior<br />

septum cannot be evaluated. The medial aspect on the right element preserves a groove<br />

that begins near the cochlear canaliculus and arches laterally to where the posterior<br />

septum meets the promontorium (i.e., where the g1 groove would also normally reach the<br />

promontorium) (Fig. 2.16A-C). There is no evidence of a foramen located on the<br />

cochlear canaliculus and leading into the promontory on this side. The groove thus has<br />

the anatomical relationships of the g4 groove of other specimens. This is the groove<br />

referred to by MacPhee et al. (1983) as the “s1” groove. They suggest it held fibers of<br />

the tympanic plexus of nerves. However, as noted above, this morphology is referred to<br />

USNM 309902 in their figure caption. The left side is differently configured. The<br />

groove and foramen relating to the tympanic canaliculus are present on the septum<br />

housing the cochlear canaliculus (Fig. 2.16A-C). However, the more ventromedial aspect<br />

of the promontorium is free of any marks. That is there is no groove that matches<br />

MacPhee et al.’s “s1” from the other side. Ventral to the fenestra vestibuli at the base of<br />

56

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