Boyer diss 2009 1046..

Boyer diss 2009 1046.. Boyer diss 2009 1046..

pages.nycep.org
from pages.nycep.org More from this publisher
01.09.2014 Views

Its anteroposterior length is 4.49 mm, while its mediolateral width is 4.11 mm. Due to broken anterior and medial margins and a missing entoglenoid process, these values are likely to underestimate the dimensions of the pristine element. The postglenoid process projects straight ventrally by 1.43 mm. On the left side, some of the zygomatic process of the squamosal is preserved (Fig. 2.8), but not enough to warrant description. Petrosal.— Both petrosals are preserved, including regions of pars cochlearis and canalicularis (Figs. 2.9-13). Remains of both ectotympanics and a substantial portion of the left auditory bulla also exist. The maximum diameter of the aperture for the fenestra vestibuli is 1.16-1.19 mm (left-right). The left spiral cochlea measures 14.5 mm in length (Fig. 2.9H) and completes 2.5 turns. The width of the pars cochlearis is 3.6-3.8 mm, while its dorsoventral depth as measured from the endocranial surface is 3.5-3.6 mm. The promontorium has only two visible grooves on its surface (Figs. 2.9-11). The identification of some of these requires consideration of how they relate to the preserved bony septa that buttress the promonotory. Four bony septa are preserved. The left promontorium visibly preserves the posterior septum (Figs. 2.10, 11) and the s1 (Fig. 2.11A-C, C’, E). HRxCT data shows that it additionally preserves the s2, and cochlear canaliculus, but no s3, arising from the medial tympanic process (Fig. 2.11E). This medial process appears to have a smooth, edge. It projects medially from the promontorium an average distance of 1.2 mm. The right promontorium preserves the cochlear canaliculus, but is broken on the medial side more anteriorly (Fig. 2.9). The g1 groove is present on the ventrolateral aspect of the promontorium. It actually grooves the ventral part of the posterior septum anteriorly; however, more posteriorly, it diverges to the lateral side of the posterior septum (Figs. 2.9-11). The g1 50

groove measures about 0.29 mm in diameter. This is the groove that MacPhee et al. (1983: fig. 1) labeled as “s2.” In their figure caption they attribute this morphology to AMNH 17388, and describe USNM 309902 as possessing the morphology of the former specimen. MacPhee et al. (1983: 509) stated that this groove has a blind end, posteriorly against a “bridge of bone uniting the promontory to the posterior wall of the bulla.” This description, however, did not reveal that the specimen is broken in some critical areas. How the bulla and promontory connected can actually only be inferred, because the whole bulla is shifted anteriorly and the posterior part of the bulla is broken out where the posterior carotid foramen would have been located (Figs. 2.9-13) on both ears. This is especially observable with the CT scan. The g2 and g5 grooves are not preserved and may or may not have been present. The g4 groove may be present on the right ear where the medial side of the promontorium is best exposed. A groove running mediolateraly on the left promontorium appears to be the g3 groove (Fig. 2.11: g3), because HRxCT data reveal that it leads towards the s2. Both ears preserve the bone-enclosed facial nerve canals (Fig. 2.9: 41). Semicircular canals are also preserved (Figs. 2.9, 11: 42). Their diameters are provided in Silcox et al. (in press). The right ear preserves a large epitympanic recess (Fig. 2.9: 43), a bone-enclosed chamber dorsolateral to the fenestra vestibuli and posterior to the epitympanic crest. The left bulla, although shifted relative to the promontorium, is fairly undistorted (Figs. 2.10-13). As indicated above, the posterior wall of the bulla is missing, however, the limestone that infills it creates a natural cast, which allows visualization of its intact 51

Its anteroposterior length is 4.49 mm, while its mediolateral width is 4.11 mm. Due to<br />

broken anterior and medial margins and a missing entoglenoid process, these values are<br />

likely to underestimate the dimensions of the pristine element. The postglenoid process<br />

projects straight ventrally by 1.43 mm. On the left side, some of the zygomatic process of<br />

the squamosal is preserved (Fig. 2.8), but not enough to warrant description.<br />

Petrosal.— Both petrosals are preserved, including regions of pars cochlearis and<br />

canalicularis (Figs. 2.9-13). Remains of both ectotympanics and a substantial portion of<br />

the left auditory bulla also exist. The maximum diameter of the aperture for the fenestra<br />

vestibuli is 1.16-1.19 mm (left-right). The left spiral cochlea measures 14.5 mm in length<br />

(Fig. 2.9H) and completes 2.5 turns. The width of the pars cochlearis is 3.6-3.8 mm,<br />

while its dorsoventral depth as measured from the endocranial surface is 3.5-3.6 mm.<br />

The promontorium has only two visible grooves on its surface (Figs. 2.9-11). The<br />

identification of some of these requires consideration of how they relate to the preserved<br />

bony septa that buttress the promonotory. Four bony septa are preserved. The left<br />

promontorium visibly preserves the posterior septum (Figs. 2.10, 11) and the s1 (Fig.<br />

2.11A-C, C’, E). HRxCT data shows that it additionally preserves the s2, and cochlear<br />

canaliculus, but no s3, arising from the medial tympanic process (Fig. 2.11E). This<br />

medial process appears to have a smooth, edge. It projects medially from the<br />

promontorium an average distance of 1.2 mm. The right promontorium preserves the<br />

cochlear canaliculus, but is broken on the medial side more anteriorly (Fig. 2.9).<br />

The g1 groove is present on the ventrolateral aspect of the promontorium. It<br />

actually grooves the ventral part of the posterior septum anteriorly; however, more<br />

posteriorly, it diverges to the lateral side of the posterior septum (Figs. 2.9-11). The g1<br />

50

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!