Boyer diss 2009 1046..

Boyer diss 2009 1046.. Boyer diss 2009 1046..

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supraoccipital are likely fused to one another on the posterior aspect of the skull because no suture separating them is evident. The dorsoventral dimensions of the supraoccipital are intact, given that the base of the nuchal crest is preserved (Figs. 2.2, 3: 31). The supraoccipital thus measures 8.30 mm dorsoventrally from the top of the foramen magnum to the top of the nuchal crest. The bone is concave in its dorsoventral profile, suggesting that the nuchal crest was prominent. Finally, the supraoccipital is marked by several small foramina. REVISED DESCRIPTION OF OTHER NORTH AMERICAN PLESIADAPID CRANIA Nannodectes intermedius USNM 309902 Despite having been figured in previous publications, most of the morphology present in this specimen remains inadequately illustrated and undescribed. Furthermore, some aspects of the petrosal description by MacPhee et al. (1983) are incorrect, due to what appears to have been an editorial mistake, as described above. In this section I revise and add to previous descriptions, assisted through HRxCT imagery. Nasal.— Remnants of both the right and left nasals are preserved (Fig. 2.8). The tip of the left element measures 2.31 mm mediolaterally (Fig. 2.8: 32). The anteroposterior length of the right counterpart measures 13.61 mm (Fig. 2.8: 33, 34). Premaxilla.— Right and left premaxillae are preserved, although the latter is extremely fragmentary (Fig. 2.8). The right side element has its dorsoventral depth (8.33 mm), palatal length (7.75 mm) and maxillary suture preserved (Fig. 2.8: 35). The dorsal 48

margin clearly had a posteriorly projecting process that most likely contacted the frontal. No teeth remain in the right premaxilla, however, Gingerich et al. (1983: fig. 5) show that both I 1 ’s and the right I 2 were preserved nearby. Lacrimal.— A fragment of the lacrimal appears to be preserved (Fig. 2.8) but none of its sutures, the lacrimal foramen or the lacrimal tubercle are observable. Maxilla.— The maxillae are preserved bilaterally. The anterior portion is broken on both sides so that the canine is missing. On the right side the P 2-3 are missing. On the left all teeth are present (Fig. 2.8) and the P 2 to M 3 distance is 12.04 mm. On the left side, the roots of the posterior molars, including M 2-3 are exposed in the orbit (Fig. 2.8: 36). None of the sutures of the maxilla are preserved except a small segment of the premaxillary suture on the right side (Fig. 2.8). The infraorbital foramen is preserved and measures 2.20 mm by 1.15 mm (Fig. 2.8: 37). Zygomatic.— A fragment of the left zygomatic bone is preserved (Fig. 2.8). It seems that most of its ventral anteroposterior length is present, measuring (9.70 mm); however the dorsal margin is broken, especially anteriorly, so that the maxillary contact is not visible. Palatines, sphenoids, frontals and parietals.— These bones may be visible, but if so, they are too fragmentary to allow certain identification or meaningful description. A delicately preserved foramen that has been pushed into the palate through deformation may represent the optic foramen, because it is similar in proportional size and morphology to optic foramina of European plesiadapid specimens (Fig. 2.8: 38). Squamosal.— The right glenoid of the squamosal (Fig. 2.9: 39) and its postglenoid process (Fig. 2.9: 40) are nearly completely preserved. The glenoid is flat. 49

margin clearly had a posteriorly projecting process that most likely contacted the frontal.<br />

No teeth remain in the right premaxilla, however, Gingerich et al. (1983: fig. 5) show that<br />

both I 1 ’s and the right I 2 were preserved nearby.<br />

Lacrimal.— A fragment of the lacrimal appears to be preserved (Fig. 2.8) but<br />

none of its sutures, the lacrimal foramen or the lacrimal tubercle are observable.<br />

Maxilla.— The maxillae are preserved bilaterally. The anterior portion is broken<br />

on both sides so that the canine is missing. On the right side the P 2-3 are missing. On the<br />

left all teeth are present (Fig. 2.8) and the P 2 to M 3 distance is 12.04 mm. On the left<br />

side, the roots of the posterior molars, including M 2-3 are exposed in the orbit (Fig. 2.8:<br />

36). None of the sutures of the maxilla are preserved except a small segment of the<br />

premaxillary suture on the right side (Fig. 2.8). The infraorbital foramen is preserved and<br />

measures 2.20 mm by 1.15 mm (Fig. 2.8: 37).<br />

Zygomatic.— A fragment of the left zygomatic bone is preserved (Fig. 2.8). It<br />

seems that most of its ventral anteroposterior length is present, measuring (9.70 mm);<br />

however the dorsal margin is broken, especially anteriorly, so that the maxillary contact<br />

is not visible.<br />

Palatines, sphenoids, frontals and parietals.— These bones may be visible, but if<br />

so, they are too fragmentary to allow certain identification or meaningful description. A<br />

delicately preserved foramen that has been pushed into the palate through deformation<br />

may represent the optic foramen, because it is similar in proportional size and<br />

morphology to optic foramina of European plesiadapid specimens (Fig. 2.8: 38).<br />

Squamosal.— The right glenoid of the squamosal (Fig. 2.9: 39) and its<br />

postglenoid process (Fig. 2.9: 40) are nearly completely preserved. The glenoid is flat.<br />

49

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