Boyer diss 2009 1046..
Boyer diss 2009 1046.. Boyer diss 2009 1046..
eveals that 1) the postgenoid process is broken off and missing, 2) a postglenoid foramen is preserved and is located medial (not posterior) to the area from which the postglenoid process would have projected (Fig. 2.2D, E: 17), and 3) there is a well-developed entoglenoid process medial to the postglenoid foramen (Fig. 2.2E: 18). Furthermore the glenoid is revealed to be rather flat, and anteroposteriorly longer (6.96 mm) than mediolaterally wide (5.38 mm). The joint surface is deformed on its medial aspect where the right M 1 or M 2 crown is pressed into it. The entoglenoid process is quite large, projecting ventrally beyond the glenoid fossa by 1.79 mm. It is oriented longitudinally, at roughly 90 degrees from the hypothetical orientation of the intact postglenoid process, and slopes medially. The width of the zygomatic process of the squamosal at its base, where it meets the glenoid, is 1.76 mm (Fig. 2.2E: 19). The length of the process, which appears to be mainly intact, is 4.95 mm. Its maximum depth is 3.77 mm. Alisphenoid and basisphenoid.—Neither of these elements is recognizably preserved. Petrosal.—The main structures in the basicranial region that are well preserved and can, therefore, be described in detail are the promontoria of the petrosals (Figs. 2.2- 5). Two isolated petrosals of Pr. gaoi will also be described here (Figs. 2.6-7). Accordingly, several major points regarding the morphology will be addressed. First, the promontorium itself will be described (its shape and the presence, position, and relationships of any grooves and foramina). Next, the morphology of the bones surrounding the promontorium will be addressed, inasmuch as this morphology relates to anatomical interpretations of previously mentioned promontorial features. The form, number and position of bony processes extending from the promontorium will then be 40
outlined. This includes description of any bulla-forming bones and bony bulla-butressing septa extending from the promontorium. The description of the promontorium will be followed by comments on the pars canalicularis of the petrosal and finally the ectotympanic bone. Table 2.3 provides a summary of morphology exhibited/preserved by each specimen, including other plesiadapid taxa to be discussed later. On UALVP 46685, crushing and shifting of the skull has obliterated most of the sutures between the petrosals and other elements (Fig. 2.2). The left petrosal is best exposed ventromedially (Figs. 2.2, 4). The pars cochlearis is visible as a fairly smooth, bulging piece of bone. It is marked by two major sets of grooves. One emerges from the posterolateral aspect, crosses the promontorium and disappears at the anteromedial aspect of the element. A groove with this particular course is recognizeable in most plesiadapid specimens (see below). It is termed the “g3” groove in further descriptions and discussions (Table 2.3; Fig. 2.4: g3). More specifically, this groove (1) stems from the area that is anterolateral to the oval window, (2) is often connected to the other set of grooves (termed “g4” here - see below) visible in this specimen and (3) leads towards a consistently present septum termed “s2” here (see below). Admittedly, to where this groove leads and how it terminates are unknown because this part of the course is not preserved in any plesiadapid specimen. The other groove marks the medial side of the promontorium and has a ventrolateral course that brings it into close proximity with the posterolateral beginning of g3, although they do not visibly meet in this particular specimen. Many other plesiadapid specimens also frequently display sets of grooves with a course similar to that just described and are referred to as “g4” grooves from here on out (Fig. 2.4: g4). In 41
- Page 17 and 18: Description . . . . . . . . . . . .
- Page 19 and 20: Craniodental material of Plesiadapi
- Page 21 and 22: Figure 2.25. MNHN CR 965 Plesiadapi
- Page 23 and 24: Figure 4.24. UM 87990 Plesiadapis c
- Page 25 and 26: Table List Chapter 2 Table 2.1. Num
- Page 27 and 28: Table 4.30. Caudal vertebrae measur
- Page 29 and 30: CHAPTER 1: INTRODUCTION Plesiadapif
- Page 31 and 32: primates, sharing many dental featu
- Page 33 and 34: were acquired. Specifically, the
- Page 35 and 36: of the plesiadapiform skeleton. Ext
- Page 37 and 38: Gervais, M.P., 1877. Enumération d
- Page 39 and 40: Szalay, F.S., 1972. Cranial morphol
- Page 41 and 42: CHAPTER 2: A REEVALUATION OF CRANIA
- Page 43 and 44: Information on the cranium of basal
- Page 45 and 46: petrosal bulla predicts that a sutu
- Page 47 and 48: History of descriptive study of ple
- Page 49 and 50: Gingerich (1971) rebutted Szalay (1
- Page 51 and 52: 9c, Gingerich (1976) labeled a groo
- Page 53 and 54: portion of the bulla medial to the
- Page 55 and 56: Bloch and Silcox (2006) described t
- Page 57 and 58: MATERIALS AND METHODS Material exam
- Page 59 and 60: whitening, dark and light areas on
- Page 61 and 62: SYSTEMATIC PALEONTOLOGY Class MAMMA
- Page 63 and 64: efore meeting a large, anteroposter
- Page 65 and 66: The canine is a simple, single-root
- Page 67: existence and/or nature of contacts
- Page 71 and 72: eneath it while also extending post
- Page 73 and 74: y a pair of parallel grooves (Fig.
- Page 75 and 76: the skull (Fig. 2.1). The length of
- Page 77 and 78: margin clearly had a posteriorly pr
- Page 79 and 80: groove measures about 0.29 mm in di
- Page 81 and 82: 2.13: 50). The suture with the supr
- Page 83 and 84: preserved (Fig. 2.15: 55). In fact,
- Page 85 and 86: the posterior septum, a deeply inci
- Page 87 and 88: Plesiadapis tricuspidens MNHN CR 12
- Page 89 and 90: is roughly 2.8 mm long. Medial to t
- Page 91 and 92: is not convoluted like many other s
- Page 93 and 94: 110) and the only squamosal/alisphe
- Page 95 and 96: two regions for the internal jugula
- Page 97 and 98: ventral to the sinuous suture (132)
- Page 99 and 100: provides measurements of these and
- Page 101 and 102: of the promontorium of the Pellouin
- Page 103 and 104: seen on the HRxCT scan is expressed
- Page 105 and 106: primates, as well as treeshrews and
- Page 107 and 108: canaliculus are present on the sept
- Page 109 and 110: it is missing from the other side o
- Page 111 and 112: observations and interpretations ma
- Page 113 and 114: REFERENCES Beard, K.C., 1993. Phylo
- Page 115 and 116: Russell, D.E., 1959. Le crâne de P
- Page 117 and 118: TABLES Table 2.1. Numerical list of
eveals that 1) the postgenoid process is broken off and missing, 2) a postglenoid foramen<br />
is preserved and is located medial (not posterior) to the area from which the postglenoid<br />
process would have projected (Fig. 2.2D, E: 17), and 3) there is a well-developed<br />
entoglenoid process medial to the postglenoid foramen (Fig. 2.2E: 18). Furthermore the<br />
glenoid is revealed to be rather flat, and anteroposteriorly longer (6.96 mm) than<br />
mediolaterally wide (5.38 mm). The joint surface is deformed on its medial aspect where<br />
the right M 1 or M 2 crown is pressed into it. The entoglenoid process is quite large,<br />
projecting ventrally beyond the glenoid fossa by 1.79 mm. It is oriented longitudinally,<br />
at roughly 90 degrees from the hypothetical orientation of the intact postglenoid process,<br />
and slopes medially. The width of the zygomatic process of the squamosal at its base,<br />
where it meets the glenoid, is 1.76 mm (Fig. 2.2E: 19). The length of the process, which<br />
appears to be mainly intact, is 4.95 mm. Its maximum depth is 3.77 mm.<br />
Alisphenoid and basisphenoid.—Neither of these elements is recognizably<br />
preserved.<br />
Petrosal.—The main structures in the basicranial region that are well preserved<br />
and can, therefore, be described in detail are the promontoria of the petrosals (Figs. 2.2-<br />
5). Two isolated petrosals of Pr. gaoi will also be described here (Figs. 2.6-7).<br />
Accordingly, several major points regarding the morphology will be addressed. First, the<br />
promontorium itself will be described (its shape and the presence, position, and<br />
relationships of any grooves and foramina). Next, the morphology of the bones<br />
surrounding the promontorium will be addressed, inasmuch as this morphology relates to<br />
anatomical interpretations of previously mentioned promontorial features. The form,<br />
number and position of bony processes extending from the promontorium will then be<br />
40