Boyer diss 2009 1046..

Boyer diss 2009 1046.. Boyer diss 2009 1046..

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eveals that 1) the postgenoid process is broken off and missing, 2) a postglenoid foramen is preserved and is located medial (not posterior) to the area from which the postglenoid process would have projected (Fig. 2.2D, E: 17), and 3) there is a well-developed entoglenoid process medial to the postglenoid foramen (Fig. 2.2E: 18). Furthermore the glenoid is revealed to be rather flat, and anteroposteriorly longer (6.96 mm) than mediolaterally wide (5.38 mm). The joint surface is deformed on its medial aspect where the right M 1 or M 2 crown is pressed into it. The entoglenoid process is quite large, projecting ventrally beyond the glenoid fossa by 1.79 mm. It is oriented longitudinally, at roughly 90 degrees from the hypothetical orientation of the intact postglenoid process, and slopes medially. The width of the zygomatic process of the squamosal at its base, where it meets the glenoid, is 1.76 mm (Fig. 2.2E: 19). The length of the process, which appears to be mainly intact, is 4.95 mm. Its maximum depth is 3.77 mm. Alisphenoid and basisphenoid.—Neither of these elements is recognizably preserved. Petrosal.—The main structures in the basicranial region that are well preserved and can, therefore, be described in detail are the promontoria of the petrosals (Figs. 2.2- 5). Two isolated petrosals of Pr. gaoi will also be described here (Figs. 2.6-7). Accordingly, several major points regarding the morphology will be addressed. First, the promontorium itself will be described (its shape and the presence, position, and relationships of any grooves and foramina). Next, the morphology of the bones surrounding the promontorium will be addressed, inasmuch as this morphology relates to anatomical interpretations of previously mentioned promontorial features. The form, number and position of bony processes extending from the promontorium will then be 40

outlined. This includes description of any bulla-forming bones and bony bulla-butressing septa extending from the promontorium. The description of the promontorium will be followed by comments on the pars canalicularis of the petrosal and finally the ectotympanic bone. Table 2.3 provides a summary of morphology exhibited/preserved by each specimen, including other plesiadapid taxa to be discussed later. On UALVP 46685, crushing and shifting of the skull has obliterated most of the sutures between the petrosals and other elements (Fig. 2.2). The left petrosal is best exposed ventromedially (Figs. 2.2, 4). The pars cochlearis is visible as a fairly smooth, bulging piece of bone. It is marked by two major sets of grooves. One emerges from the posterolateral aspect, crosses the promontorium and disappears at the anteromedial aspect of the element. A groove with this particular course is recognizeable in most plesiadapid specimens (see below). It is termed the “g3” groove in further descriptions and discussions (Table 2.3; Fig. 2.4: g3). More specifically, this groove (1) stems from the area that is anterolateral to the oval window, (2) is often connected to the other set of grooves (termed “g4” here - see below) visible in this specimen and (3) leads towards a consistently present septum termed “s2” here (see below). Admittedly, to where this groove leads and how it terminates are unknown because this part of the course is not preserved in any plesiadapid specimen. The other groove marks the medial side of the promontorium and has a ventrolateral course that brings it into close proximity with the posterolateral beginning of g3, although they do not visibly meet in this particular specimen. Many other plesiadapid specimens also frequently display sets of grooves with a course similar to that just described and are referred to as “g4” grooves from here on out (Fig. 2.4: g4). In 41

eveals that 1) the postgenoid process is broken off and missing, 2) a postglenoid foramen<br />

is preserved and is located medial (not posterior) to the area from which the postglenoid<br />

process would have projected (Fig. 2.2D, E: 17), and 3) there is a well-developed<br />

entoglenoid process medial to the postglenoid foramen (Fig. 2.2E: 18). Furthermore the<br />

glenoid is revealed to be rather flat, and anteroposteriorly longer (6.96 mm) than<br />

mediolaterally wide (5.38 mm). The joint surface is deformed on its medial aspect where<br />

the right M 1 or M 2 crown is pressed into it. The entoglenoid process is quite large,<br />

projecting ventrally beyond the glenoid fossa by 1.79 mm. It is oriented longitudinally,<br />

at roughly 90 degrees from the hypothetical orientation of the intact postglenoid process,<br />

and slopes medially. The width of the zygomatic process of the squamosal at its base,<br />

where it meets the glenoid, is 1.76 mm (Fig. 2.2E: 19). The length of the process, which<br />

appears to be mainly intact, is 4.95 mm. Its maximum depth is 3.77 mm.<br />

Alisphenoid and basisphenoid.—Neither of these elements is recognizably<br />

preserved.<br />

Petrosal.—The main structures in the basicranial region that are well preserved<br />

and can, therefore, be described in detail are the promontoria of the petrosals (Figs. 2.2-<br />

5). Two isolated petrosals of Pr. gaoi will also be described here (Figs. 2.6-7).<br />

Accordingly, several major points regarding the morphology will be addressed. First, the<br />

promontorium itself will be described (its shape and the presence, position, and<br />

relationships of any grooves and foramina). Next, the morphology of the bones<br />

surrounding the promontorium will be addressed, inasmuch as this morphology relates to<br />

anatomical interpretations of previously mentioned promontorial features. The form,<br />

number and position of bony processes extending from the promontorium will then be<br />

40

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