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Boyer diss 2009 1046..

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Outgroup taxa were scored from the literature including Bloch et al. (2001), Clemens<br />

(2004), and Silcox and Gunnell (2008) for P. janisae, and Bloch et al. (2001) and Silcox<br />

et al. (2001) for Elphidotarsius. Elphidotarsius is a composite of E. florencae and E.<br />

wightoni, the two most basal species of the Carpolestidae (Bloch et al., 2001; Silcox et<br />

al., 2001), because individual species are fragmentarily known. The 28 plesiadapid<br />

species included were scored based on Gingerich (1976), Fox (1990), newly available<br />

data from the upper dentition of Pr. gaoi (Chapter 2), an undescribed upper dentition of<br />

Ch. caesor (CM 72770), undescribed lower incisors of Ch. caesor, and undescribed<br />

central incisors of Nannodectes gazini (CM 76922, CM 76938).<br />

Gingerich (1976) inferred dental formulae for several poorly preserved<br />

plesiadapid species. I follow these inferences in my character coding, unless evidence<br />

documented by Gingerich (1976) or elsewhere (i.e., in newly available specimens)<br />

contradicts his reconstructions. For instance, Gingerich (1976) inferred that P. cookei,<br />

Pl. russelli and other Platychoerops species lacked upper canines, but retained upper<br />

second premolars, based on the well-known morphology of P. tricuspidens. However, as<br />

documented and discussed in Chapter 3, P. cookei UM 87990 actually lacks C 1 and P 2 .<br />

Furthermore, it appears to me that the type of Pl. richardsoni (YM 550) also lacks these<br />

teeth. Therefore, it cannot be assumed that Pl. russelli and Pl. daubrei retained P 2 .<br />

The character matrix was analyzed with the program Nona (Nixon, 1999-2002) in<br />

WinClada (Goloboff, 1999) using a heuristic search of ~5,000 replicates.<br />

Character optimization - Cranial and postcranial characters discussed in the<br />

preceding three chapters (2-4), as well as characters from Bloch et al. (2007), are<br />

optimized onto the ancestral node for Plesiadapidae using the newly generated species<br />

505

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