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Boyer diss 2009 1046..

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value for P. tricuspidens. In summary, it seems likely that P. tricuspidens (like P. cookei,<br />

P. insignis, and Nannodectes) will be revealed to have phalangeal indices of around 130-<br />

140 when more confidently identified digit rays are available. This is less than that of<br />

most arboreal euprimates, but much higher than that of many terrestrial mammals (Kirk<br />

et al., 2008). If, on the other hand, it turns out that P. tricuspidens truly does have such<br />

short fingers, then this information appears to contradict much other information from the<br />

skeleton suggesting arboreal habits (Youlatos and Godinot, 2004). It should, however, be<br />

noted that it would be consistent with the absolutely thicker and proportionally shorter<br />

limbs of P. tricuspidens as compared to those of P. cookei and extant arboreal mammals<br />

(Runestad and Ruff, 1995).<br />

Body mass estimates<br />

Differences in results of the two sets of postcranial regressions used to generate<br />

body mass estimates are interesting because they suggest that P. cookei has exceptionally<br />

relatively long limbs among mammals (based on the Gingerich [1990] equations), but<br />

relatively short limbs compared with extant primates (based on the unpublished extant<br />

primate sample). The absolute body mass estimates generated by both sets of postcranial<br />

regressions are of some interest because they sample the skeleton so broadly and show<br />

such congruence that they may actually be fairly accurate. That is, P. cookei probably<br />

weighed somewhere between 1-3 kg, but not as much as the 5 kg suggested by a<br />

regression from Silcox et al. (in press) based on skull length of extant primates.<br />

Proportional differences in body mass between P. cookei and other plesiadapids<br />

as suggested by the postcranium are much smaller than estimates of proportional<br />

388

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