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Boyer diss 2009 1046..

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A second analysis includes all seven measurements (Fig. 4.50B). P. insignis is<br />

not included because metapodial data are not available for it. The positions of the<br />

included plesiadapiforms are similar to what they are in the previous analysis: they plot<br />

close to each other, are surrounded by arboreal taxa, and are not particularly close to<br />

other euarchontans, Tupaia or Cynocephalus.<br />

The third analysis includes all of the measurements of the second analysis except<br />

for trunk length (Trk-L was removed from all calculations) (Fig. 4.50C). This allows the<br />

inclusion of a larger, more diverse taxon sample. The positions of the plesiadapiform<br />

taxa are similar to what they are in the other two analyses. Interestingly the primitive,<br />

arboreal treeshrew, Ptilocercus lowii, plots close to the plesiadapiforms. Measurements<br />

representing a composite micromomyid plesiadapiform were included and show it to plot<br />

close to Carpolestes and P. cookei. Interestingly cercopithecid euprimates plot in the<br />

same part of the morphospace as some of these plesiadapiforms. Unlike other extant taxa<br />

in this region of the morphospace, the sampled cercopithecids (Table 4.35) are relatively<br />

terrestrially adapted among euprimates.<br />

Comparing limb indices of P. cookei to those of other plesiadapids reveals that it<br />

has a short trunk (Table 4.34: Tr-I) compared to its limb lengths, a long radius and femur<br />

compared to its humerus length (Table 4.34: Br-I and Hf-I), a tibia that is short compared<br />

to the radius but long compared to the femur (Table 4.34: Rt-I and Cr-I), and a forelimb<br />

that is long compared to the hind limb (Table 4.34:Int-I). One exception here is that P.<br />

insignis has an even longer femur relative to its humeral length. Most of these metrics<br />

cannot be evaluated in P. tricuspidens because no tibial lengths are available (thus<br />

previous calculations of intermembral index are estimates: e.g., Gingerich, 1976).<br />

376

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