Boyer diss 2009 1046..
Boyer diss 2009 1046.. Boyer diss 2009 1046..
Manubrium and Sternebrae Description.—The manubrium and six sternebrae are preserved (Fig. 4.48; Table 4.31). None of these elements are known for any other plesiadapiforms. The manubrium is longer than any of the sternebrae (Table 4.31). The right costal process is broken but the left one is intact. Its lateral edge received the first rib. On its craniodorsal aspect a slight depression for the clavicle articulation is observable. The cranial margin of the manubrium is convex. The dorsal surface is flat. The ventral surface is marked by a strong midline keel. The keel is narrowest at its root. Thus, a transverse cross-section through the manubrium reveals it to be in the shape of an “I-beam.” The articular surface for the first sternebra is oriented somewhat ventrally. Thus, when articulated, the manubrium and first sternebra form an obtuse ventral angle. The sternebrae are blocky bones and are all fairly similar to each other. The first is strongly keeled, while the keel becomes increasingly muted on more caudal positions. The last sternebra differs from the others in having a caudal end that is rounded instead of flat, and in having a longitudinal groove on its dorsal surface. Comparison.—The manubrium of P. cookei appears more similar to that of Tupaia glis than to those of many euprimates in having a stronger keel and in forming a strong angle with the first sternebra. Ribs Description.—Twelve rib fragments are attributed to UM 87990 (Fig. 4.49; Table 4.32). Six appear to be left side elements. One or two of the floating ribs (T10-13) are 374
preserved. The ribs are slender and most exhibit a crest along their caudal borders. This crest is most expansive near the rib angles and becomes less pronounced farther ventrally. On the medial surface of these crests is a groove for the intercostal neurovasculature. Comparison.—The ribs compare best to those of tupaiid treeshrews and other plesiadapiforms (Boyer and Bloch, 2008) among euarchontan mammals. Ptilocercid treeshrews, dermopterans, and many euprimates have ribs that are much more craniocaudally expanded (Sargis, 2001) than those of P. cookei. N. gidleyi preserves rib fragments showing it to be similar to P. cookei in this regard (Table 4.32). Body proportions in P. cookei and other plesiadapids A set of seven body segment lengths were analyzed with three separate principal coordinate analyses of the Euclidean distance matrices relating cases in the sample (measurements include: Trk-L, H-L, R-L, MC III-L, F-L, T-L, and MT III-L – see Tables 4.33-35; Fig. 4.50). As discussed in the Methods section, the variables used to create the distance matrices are log ratios of each measurement to the geometric mean of all measurements for a specimen in a given analysis. The first analysis does not include metapodial length data (MC III-L and MT III-L were removed from all calculations), and can therefore include measurements of the skeleton of P. insignis, on which MT III-L is unmeasurable (Gingerich, 1976) (Fig. 4.50A). It shows P. insignis to plot by itself, closest to terrestrial and scansorial rodents and Tupaia glis. P. cookei plots nearby, but is in the middle of a distribution including various arboreal taxa. Other plesiadapiforms, Ignacius and Carpolestes plot even farther from P. insignis. Cynocephalus volans plots far from all of these taxa. 375
- Page 351 and 352: antipronograde clinging postures, o
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Manubrium and Sternebrae<br />
Description.—The manubrium and six sternebrae are preserved (Fig. 4.48; Table<br />
4.31). None of these elements are known for any other plesiadapiforms. The manubrium<br />
is longer than any of the sternebrae (Table 4.31). The right costal process is broken but<br />
the left one is intact. Its lateral edge received the first rib. On its craniodorsal aspect a<br />
slight depression for the clavicle articulation is observable. The cranial margin of the<br />
manubrium is convex. The dorsal surface is flat. The ventral surface is marked by a<br />
strong midline keel. The keel is narrowest at its root. Thus, a transverse cross-section<br />
through the manubrium reveals it to be in the shape of an “I-beam.”<br />
The articular surface for the first sternebra is oriented somewhat ventrally. Thus,<br />
when articulated, the manubrium and first sternebra form an obtuse ventral angle. The<br />
sternebrae are blocky bones and are all fairly similar to each other. The first is strongly<br />
keeled, while the keel becomes increasingly muted on more caudal positions. The last<br />
sternebra differs from the others in having a caudal end that is rounded instead of flat,<br />
and in having a longitudinal groove on its dorsal surface.<br />
Comparison.—The manubrium of P. cookei appears more similar to that of<br />
Tupaia glis than to those of many euprimates in having a stronger keel and in forming a<br />
strong angle with the first sternebra.<br />
Ribs<br />
Description.—Twelve rib fragments are attributed to UM 87990 (Fig. 4.49; Table<br />
4.32). Six appear to be left side elements. One or two of the floating ribs (T10-13) are<br />
374