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Boyer diss 2009 1046..

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Caudal vertebrae<br />

Description.—There are 17 caudal (Ca) vertebrae attributed to UM 87990 (Fig.<br />

4.45; Table 4.30). The most proximal position preserved (Fig. 4.45A) has a complete<br />

vertebral canal, pre- and postzygapophyses, posteriorly projecting transverse processes,<br />

and the broken base of a spinous process. The centrum is missing but it would have been<br />

relatively craniocaudally short and mediolaterally wide. Typically there are three<br />

proximal caudal vertebrae with these basic proportions, as there are in the most<br />

completely known plesiadapiform tail of Ignacius clarkforkensis (<strong>Boyer</strong> and Bloch,<br />

2008). The next most proximal caudal vertebra still retains a complete vertebral canal,<br />

but it is substantially more elongate than the first (but shorter and less elongate than the<br />

next in the series). This appears to be either Ca4 or Ca5 (Fig. 4.45B). The following<br />

vertebra appears to be Ca6 (Fig. 4.45C). It is much longer than the first two. It has an<br />

incomplete vertebral canal (the laminae do not meet in the sagittal plane). The pre- and<br />

post zygapophyses did not directly articulate with those of more proximal and distal<br />

vertebrae, respectively. There is a set of cranioventrally projecting transverse processes<br />

and a set of caudoventrally projecting processes. The vertebra identified as Ca7 is very<br />

similar to Ca6. It differs in being absolutely longer, proportionally narrower, and in<br />

having absolutely and proportionally smaller transverse processes. From Ca8-Ca12 (Fig.<br />

4.45E-I) the vertebrae are identified on the basis of increasing length and/or decreasing<br />

prominence of zygapophyses and transverse processes. The centrum diameters of these<br />

vertebrae do not appear to vary drastically or systematically through this region. Ca11<br />

and Ca12 (Fig. 4.45H, I) are extremely similar to one another and it is possible that their<br />

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