Boyer diss 2009 1046..
Boyer diss 2009 1046.. Boyer diss 2009 1046..
including the lumbar, sacral and proximal part of the caudal region. Generally speaking, vertebrae described in this section were identified as being part of the thoracic region by the presence of rib facets on the body or transverse processes and in some cases by the orientation of the zygapophyseal facets. The craniocaudal position of a given thoracic vertebra, other than whether it comes before or after the diaphragmatic position is difficult to determine due to extensive postmortem deformation that prevents assessments through articulation or detailed measurements. Often centrum length, height or overall size increases progressively from anterior to posterior. The first thoracic vertebra (Fig. 4.42A) is craniocaudally shorter than the others, but there is no pattern of increase in the remaining prediaphragmatic vertebrae (Table 4.27; Fig. 4.42B-I). This is demonstrated beyond any doubt by the only two vertebrae preserved in articulation (Fig. 4.42E), in which the more cranial one is slightly, but definitely, longer. However, the two postdiaphragmatic thoracic vertebrae (Fig. 4.42J-K) are longer than the ten vertebrae cranial to them. The first three thoracic vertebrae exhibit dorsoventrally shallow cranial and caudal centrum facets, while T4-T10 are deeper, with no discernable pattern of increase or decrease. T12 cannot be measured cranially and T13 cannot be measured caudally, but together they are the deepest caudally and deepest cranially (respectively) in the thoracic region. Centrum mediolateral widths are even less tractable. T1 is wider cranially than all vertebrae that follow until T13. Caudally it is the widest of the thoracic region. Of the vertebrae with preserved transverse processes, the most caudal ones are mediolaterally narrower than the more anterior ones (i.e., Fig. 4.42E vs. 4.42H-I). Of those with intact laminae and spinous processes, the more posterior postions have more vertically oriented processes. The prediaphragmatic spinous processes are all longer 366
dorsoventrally than craniocaudally. The thoracic vertebrae have zygapophyses with rounded margins, although T12 appears to have postzygapophyses that are slightly more angular, or square. The postzygapophyses of T13 are too broken for meaningful comment. The postdiaphragmatic spinous process of T12 is broken, but was probably vertically oriented, revealing it as the anticlinal vertebra. The T13 process is badly broken, but its caudal and dorsocaudal margins are intact, revealing that it was cranially oriented and most likely shorter dorsoventrally than craniocaudally long. The two postdiaphragmatic vertebrae preserve the roots of large accessory processes (anapophyses). Comparison.—To have 13 thoracic vertebrae is slightly above average among small generalized, terrestrial to arboreal mammals (Table 4.24). It is the same number seen in tupaiid treeshrews (13), but one less than in Ptilocercus and Cynocephalus (14) (Sargis, 2001). It is generally thought that a longer thorax represents an emphasis on stabilization rather than mobility (Sargis, 2001). The presence of an anticlinal vertebra within the thoracic region, and large anapophyses, indicates that this region had pronounced mobility in dorsiflexion and extension, but was restricted in axial rotation or medial flexion. Animals that sometimes use a bounding, asymmetrical gait, and which therefore do not require dorsostability, are charactized by such features (e.g., Sciurus, treeshrews, many arboreal quadrupedal primates). Apes, humans, lorises, sloths, Cynocephalus and some opossums lack a thoracic anticlinal vertebra, and do not engage in quadrupedal bounding gaits (Boyer and Bloch, 2008). The thoracic region of P. cookei therefore indicates that it was not restricted to slow, cautious locomotion. 367
- Page 343 and 344: even more pronounced. The distal en
- Page 345 and 346: etween the distal carpals and the
- Page 347 and 348: have stouter shaft diameters for th
- Page 349 and 350: difference makes them more like kno
- Page 351 and 352: antipronograde clinging postures, o
- Page 353 and 354: foramina, and faces slightly proxim
- Page 355 and 356: spine at the superior tip of the il
- Page 357 and 358: the thigh (Gambaryan, 1974). The ha
- Page 359 and 360: The femoral shaft is smooth, lackin
- Page 361 and 362: lacking a lateral extension of its
- Page 363 and 364: The anteromedial side of the tibial
- Page 365 and 366: fibular notch and the strong crest
- Page 367 and 368: to the peroneal surface. The perone
- Page 369 and 370: could even be described as having t
- Page 371 and 372: Function.—The functional features
- Page 373 and 374: flexor fibularis groove surface. Ex
- Page 375 and 376: tubercle, which is centrally locate
- Page 377 and 378: Cuboid Description.—The right cub
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- Page 399 and 400: vertebrae. It is also similar to th
- Page 401 and 402: identifications have been reversed.
- Page 403 and 404: preserved. The ribs are slender and
- Page 405 and 406: Carpolestes simpsoni (UM 101963) an
- Page 407 and 408: third metacarpal, similar to arbore
- Page 409 and 410: autapomorphy, because it appears th
- Page 411 and 412: Jenkins (1974) found that Tupaia gl
- Page 413 and 414: Nannodectes and other plesiadapids,
- Page 415 and 416: are consistent with more frequent u
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- Page 419 and 420: Perry, M. Silcox, R. Secord and man
- Page 421 and 422: ubriventer: implications for the fu
- Page 423 and 424: Linnaeus, C., 1758. Systema naturae
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- Page 427 and 428: Table 4.3A. Measurements of the pro
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including the lumbar, sacral and proximal part of the caudal region. Generally speaking,<br />
vertebrae described in this section were identified as being part of the thoracic region by<br />
the presence of rib facets on the body or transverse processes and in some cases by the<br />
orientation of the zygapophyseal facets. The craniocaudal position of a given thoracic<br />
vertebra, other than whether it comes before or after the diaphragmatic position is<br />
difficult to determine due to extensive postmortem deformation that prevents assessments<br />
through articulation or detailed measurements. Often centrum length, height or overall<br />
size increases progressively from anterior to posterior. The first thoracic vertebra (Fig.<br />
4.42A) is craniocaudally shorter than the others, but there is no pattern of increase in the<br />
remaining prediaphragmatic vertebrae (Table 4.27; Fig. 4.42B-I). This is demonstrated<br />
beyond any doubt by the only two vertebrae preserved in articulation (Fig. 4.42E), in<br />
which the more cranial one is slightly, but definitely, longer. However, the two<br />
postdiaphragmatic thoracic vertebrae (Fig. 4.42J-K) are longer than the ten vertebrae<br />
cranial to them. The first three thoracic vertebrae exhibit dorsoventrally shallow cranial<br />
and caudal centrum facets, while T4-T10 are deeper, with no discernable pattern of<br />
increase or decrease. T12 cannot be measured cranially and T13 cannot be measured<br />
caudally, but together they are the deepest caudally and deepest cranially (respectively) in<br />
the thoracic region. Centrum mediolateral widths are even less tractable. T1 is wider<br />
cranially than all vertebrae that follow until T13. Caudally it is the widest of the thoracic<br />
region. Of the vertebrae with preserved transverse processes, the most caudal ones are<br />
mediolaterally narrower than the more anterior ones (i.e., Fig. 4.42E vs. 4.42H-I). Of<br />
those with intact laminae and spinous processes, the more posterior postions have more<br />
vertically oriented processes. The prediaphragmatic spinous processes are all longer<br />
366