Boyer diss 2009 1046..
Boyer diss 2009 1046.. Boyer diss 2009 1046..
face ventrally. The centrum is rectangular with a distinct midline ventral ridge. Its cranial facet faces cranially and is angled slightly ventrally. Within the vertebral canal, the dorsal surface of the centrum is marked by two foramina. The transverse processes are long and narrow, projecting inferolaterally. Another cervical vertebra is represented by the centrum alone (Fig. 4.41D). It is considered to be C4, even though it cannot be articulated with C3, due to the missing caudal C3 epiphysis. There is slight mismatch between the caudal epiphysis of “C4” and C6, which is taken as additional evidence that it is correctly identified as C4. The C4 centrum is even more “rectangular” than that of C3, because the dorsal and ventral surfaces are parallel to one another, and there is no midline ridge on the ventral surface. Like C3, there are two foramina on the dorsal surface. The cranial centrum facet faces slightly ventrally, like that of C3. The caudal facet faces slightly dorsally; however it has a dorsal lip, which makes the surface cylindrically concave. The last preserved cervical vertebra is identified as C6, mainly due to the presence of broken roots of hypertrophied anterior tubercles, as typically characterize this position (Fig. 4.41E). The left half of the pedicle and lamina are also broken away, as are the tips of the transverse processes. The prezygapophyses are oriented slightly more medially than the dorsally facing facets of C3, while the postzygapophyseal facets are oriented slightly more laterally than the ventrally facing facets of C3. The foramina transversaria are preserved. The centrum is basically identical in shape and morphology to that of C4. Comparison.—Again, a comparative context is needed to comment on functionally salient features of the cervical vertebrae. C3, C4 and C6 in P. cookei have morphology similar to that preserved in N. gidleyi and N. intermedius, although these 364
taxa appear to have slightly more prominent ventral midline ridges on their vertebrae. Plesiadapids are also morphologically similar to Ptilocercus and arboreal tree squirrels in (1) having vertebral bodies of the cervical vertebrae that are mediolaterally wide relative to their craniocaudal length and (2) lacking pronounced spinous processes. On the other hand, Tupaia and terrestrial rodents (e.g., Rattus) have mediolaterally narrower vertebral bodies and more pronounced spinous processes. These differences have been related to contrasts in head mobility among various taxa (Argot, 2002). The tupaiid-like form is thought to retain greater mediolateral mobility due to the presence of a narrower body. Cynocephalus is an interesting case because it has cervical centra that are similar to those of plesiadapids in most respects, but it has prominent spinous processes on all of its cervical vertebrae, unlike plesiadapids (Stafford, 1999). Table 4.24 gives neck length as a proportion of trunk length. P. cookei is similar to most of the relatively small scansorial-to-arboreal taxa in its proportional neck length. Cynocephalus and Tarsius have much longer necks. As stated above, principal coordinate analyses also appear to differentiate these two taxa on the basis of their long necks. Stafford (1999) presents data showing Cynocephalus, bats and sloths to have exceptionally long necks among arboreal to scansorial mammals. Thoracic vertebrae Description.—All but one (12 out of 13) of the thoracic (T) vertebrae appear to be preserved (Fig. 4.42). The missing position is the diaphragmatic vertebra, which should have dorsolaterally angled prezygapophyseal facets, like the vertebrae more cranial to it, and dorsomedially facing postzygapophyseal facets, like the vertebrae more caudal to it, 365
- Page 341 and 342: differs from MC II and III in havin
- Page 343 and 344: even more pronounced. The distal en
- Page 345 and 346: etween the distal carpals and the
- Page 347 and 348: have stouter shaft diameters for th
- Page 349 and 350: difference makes them more like kno
- Page 351 and 352: antipronograde clinging postures, o
- Page 353 and 354: foramina, and faces slightly proxim
- Page 355 and 356: spine at the superior tip of the il
- Page 357 and 358: the thigh (Gambaryan, 1974). The ha
- Page 359 and 360: The femoral shaft is smooth, lackin
- Page 361 and 362: lacking a lateral extension of its
- Page 363 and 364: The anteromedial side of the tibial
- Page 365 and 366: fibular notch and the strong crest
- Page 367 and 368: to the peroneal surface. The perone
- Page 369 and 370: could even be described as having t
- Page 371 and 372: Function.—The functional features
- Page 373 and 374: flexor fibularis groove surface. Ex
- Page 375 and 376: tubercle, which is centrally locate
- Page 377 and 378: Cuboid Description.—The right cub
- Page 379 and 380: Ectocuneiform Description.—A left
- Page 381 and 382: Metatarsals Hallucal metatarsal des
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- Page 385 and 386: medial side facet on MT IV from a v
- Page 387 and 388: Vertebral column Vertebral column d
- Page 389 and 390: measurements, see caption of Fig. 4
- Page 391: transverse processes, and the poste
- Page 395 and 396: dorsoventrally than craniocaudally.
- Page 397 and 398: The zygapophyses increase in size b
- Page 399 and 400: vertebrae. It is also similar to th
- Page 401 and 402: identifications have been reversed.
- Page 403 and 404: preserved. The ribs are slender and
- Page 405 and 406: Carpolestes simpsoni (UM 101963) an
- Page 407 and 408: third metacarpal, similar to arbore
- Page 409 and 410: autapomorphy, because it appears th
- Page 411 and 412: Jenkins (1974) found that Tupaia gl
- Page 413 and 414: Nannodectes and other plesiadapids,
- Page 415 and 416: are consistent with more frequent u
- Page 417 and 418: differences generated from informat
- Page 419 and 420: Perry, M. Silcox, R. Secord and man
- Page 421 and 422: ubriventer: implications for the fu
- Page 423 and 424: Linnaeus, C., 1758. Systema naturae
- Page 425 and 426: Stern, J.T.,Jr., 1988. Essentials o
- Page 427 and 428: Table 4.3A. Measurements of the pro
- Page 429 and 430: Table 4.3C. Comparative shape varia
- Page 431 and 432: Table 4.5A. Measurements and shape
- Page 433 and 434: Table 4.10. Measurements and shape
- Page 435 and 436: Table 4.13. Measurements of plesiad
- Page 437 and 438: Table 4.15. Measurements of plesiad
- Page 439 and 440: Table 4.17B. Measurements of the di
- Page 441 and 442: Table 4.18B. Measurements of the di
face ventrally. The centrum is rectangular with a distinct midline ventral ridge. Its<br />
cranial facet faces cranially and is angled slightly ventrally. Within the vertebral canal,<br />
the dorsal surface of the centrum is marked by two foramina. The transverse processes<br />
are long and narrow, projecting inferolaterally.<br />
Another cervical vertebra is represented by the centrum alone (Fig. 4.41D). It is<br />
considered to be C4, even though it cannot be articulated with C3, due to the missing<br />
caudal C3 epiphysis. There is slight mismatch between the caudal epiphysis of “C4” and<br />
C6, which is taken as additional evidence that it is correctly identified as C4. The C4<br />
centrum is even more “rectangular” than that of C3, because the dorsal and ventral<br />
surfaces are parallel to one another, and there is no midline ridge on the ventral surface.<br />
Like C3, there are two foramina on the dorsal surface. The cranial centrum facet faces<br />
slightly ventrally, like that of C3. The caudal facet faces slightly dorsally; however it has<br />
a dorsal lip, which makes the surface cylindrically concave.<br />
The last preserved cervical vertebra is identified as C6, mainly due to the presence<br />
of broken roots of hypertrophied anterior tubercles, as typically characterize this position<br />
(Fig. 4.41E). The left half of the pedicle and lamina are also broken away, as are the tips<br />
of the transverse processes. The prezygapophyses are oriented slightly more medially<br />
than the dorsally facing facets of C3, while the postzygapophyseal facets are oriented<br />
slightly more laterally than the ventrally facing facets of C3. The foramina transversaria<br />
are preserved. The centrum is basically identical in shape and morphology to that of C4.<br />
Comparison.—Again, a comparative context is needed to comment on<br />
functionally salient features of the cervical vertebrae. C3, C4 and C6 in P. cookei have<br />
morphology similar to that preserved in N. gidleyi and N. intermedius, although these<br />
364