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Boyer diss 2009 1046..

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lateral trochleae is usually larger, or projects farther distally than the other. Typically this<br />

larger trochlea is on the opposite side of that sporting the larger, blunt proximal tubercle.<br />

As can be seen in lateral and dorsal views, the distal articular surface has its largest radius<br />

of curvature on its ventral aspect, a narrower one distally, and almost no expression of the<br />

distal articular surface on its dorsal aspect. The lateral aspects of the distal end are pitted<br />

for collateral ligaments.<br />

Lateral proximal phalanx function.—The shape of the proximal articular surface<br />

of the proximal phalanx and its orientation with respect to the shaft, combined with<br />

articulating metacarpals in which the distal articular surfaces are spherical and oriented<br />

dorsally, suggest a metacarpophalangeal joint that was (1) mobile in abduction-adduction,<br />

axial rotation, and flexion-extension; and (2) was most stable in a dorsiflexed position,<br />

where the greatest overlap in corresponding joint surface area is formed. The orientation<br />

of the distal articular surface of the proximal phalanx, its differential radius of curvature,<br />

and the presence of extensive flexor sheath ridges suggest habitual ventroflexion of the<br />

proximal interphalangeal joints.<br />

Lateral proximal phalanges comparions.—Compared to other plesiadapids, P.<br />

cookei cannot be demonstrated to differ strongly in any easily quantifiable fashion.<br />

Smaller Nannodectes species seem to exhibit less pronounced lateral trochleae on the<br />

distal articular surfaces and to have proportionally slightly wider shafts. Some of the<br />

proximal phalanges from the French Berru locality have prominent ventrally projecting<br />

flexor sheath ridges (e.g., MNHN R 503; MNHN unnumbered Divers coll. specimen: see<br />

Table 4.13). Although they are identified as plesidapids, this noted morphologic<br />

320

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