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Boyer diss 2009 1046..

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Function.—The three proximal carpal articular surfaces of the hamate suggest it to<br />

have been a highly mobile bone (along with the rest of the distal carpal row) that was<br />

capable of taking on many different positions and articular configurations with respect to<br />

the rest of the carpus.<br />

Comparison.—Compared to the hamates known for P. tricuspidens and N.<br />

intermedius, that of P. cookei differs in several respects. The radioulnar width of the<br />

distal end in the hamate of P. cookei is narrower relative to its dorsoventral depth (Table<br />

4.9: DEV). The capitate articular surface is concave instead of flat. Finally, the proximal<br />

margin has a more distinctive facet for the scaphoid than seen in either N. intermedius or<br />

P. tricuspidens (Fig. 4.13B: 3). The greater development of the proximal scaphoid facet<br />

may suggest more frequent reliance on adducted hand postures, because in order for this<br />

facet to contact the scaphoid solidly the hamate must be adducted with respect to it.<br />

Although the differences between P. cookei and the other plesidapids do not seem great,<br />

they are not trivial either. In the absence of better cranial-postcranial associations for the<br />

hamate of UM 87990, it is at least possible that these differences indicate that this bone<br />

does not actually belong to P. cookei.<br />

Metacarpals<br />

There are two different MC II-V morphs that occur with the specimen (set 1 - Fig.<br />

4.14; sets 1-2 – Fig. 4.15; set 2 - Fig. 4.16; Tables. 4.10-12). The two sets are of similar<br />

size and morphology. It is difficult to determine which elements belong to which set and<br />

further which belong to P. cookei. Comparisons to other better-associated plesiadapid<br />

307

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