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Boyer diss 2009 1046..

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carpals (Figs. 4.7B, 4.9A: 2). Additionally, the shelf-like distally facing facet on the<br />

scaphoid continues ulnarly with morphology of the lunate (Figs. 4.7B, 4.9A: 4). The<br />

radius articular surface of the lunate is also visible in dorsal view (Figs. 4.7B, 4.9A: 6). It<br />

arcs from the radiodistal edge of the lunate, down to the lunate’s proximoulnar margin,<br />

where it contacts the dorsal edge of a small, ulnar facing facet for the distal part of the<br />

distal radial facet of the ulna (Figs. 4.7B, 4.9A: 3). This facet for the ulna can be seen<br />

more clearly from ulnar view. This view also shows that the lunate is mainly nonarticular<br />

on the ulnar side. Its proximal surface has the facet for the ulna. Distal to this<br />

there is a non-articular gap, and farther distal still there is a strip-like facet for the<br />

triquetrum, which faces laterodistally (Fig. 4.7B, 4.9A: 1).<br />

Function.—The morphology of the lunate, like that of the scaphoid, suggests a<br />

habitually dorsiflexed carpus, and the capacity for a large amount of dorsiflexion.<br />

Comparison.—Comparing the lunate described here to that described for N.<br />

intermedius by Beard (1989) (Fig. 4.9A, C), one must conclude that the two bones are<br />

very different. This is surprising given (1) the similarity of the scaphoids of these two<br />

taxa (Fig. 4.8) and (2) the good fit between the scaphoid and the lunate described here.<br />

These two facts begin to suggest that the bone described as the lunate in N. intermedius is<br />

probably incorrectly identified as such, or incorrectly attributed to N. intermedius. Two<br />

observations support the former hypothesis. A bone with nearly identical morphology<br />

was prepared while preserving its association with an intermediate phalanx of USNM<br />

442229 (Fig. 4.9D), strongly suggesting the bone identified by Beard as the lunate does<br />

in fact belong to N. intermedius. However, if this bone is actually the lunate, then the<br />

newly identified element and the originally described one should represent opposite sides<br />

300

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