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Boyer diss 2009 1046..

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distinct, convex distal radial facet reinforces the inference that there was substantial axial<br />

mobility in the radius with respect to the ulna. Its location on the dorsal aspect of the ulna<br />

is consistent with morphology of the radius, suggesting that the wrist was maintained in a<br />

semi-supinated orientation most of the time.<br />

Comparison.—In general, the ulna of P. cookei does not differ appreciably from<br />

those of other plesiadapids. None of the shape indices in Table 4.6A reveal any<br />

quantified differences. Qualitatively comparing more subtle aspects of the morphology,<br />

it can be stated that the ulna of P. cookei seems to be unique in the shallowness of the<br />

longitudinal groove marking the lateral side of its shaft for origination of extensor<br />

musculature. The ulna of P. cookei also appears more gracile than those of other<br />

plesiadapids; however, as mentioned, this is not borne out by any measurements (e.g.,<br />

Table 4.5A: NSV and SSV).<br />

Scaphoid<br />

Description.—The right scaphoid is preserved (Table 4.6; Fig. 4.7A). It was<br />

originally identified by comparisons to the scaphoid of N. intermedius USNM 442229<br />

(Beard, 1993a) and a scaphoid dentally associated to the paromomyid Acidomomys<br />

hebeticus (<strong>Boyer</strong> and Bloch, 2008). Aside from being “boat-shaped,” as the etymology<br />

of the bone’s name suggests, with a convex proximal articular surface forming the “hull<br />

of the boat” and the concave distal articular surface forming the “boat’s interior,” the<br />

scaphoid of P. cookei could also be described as “barbell-shaped” because the ventral<br />

tubercle is quite large and distinct from the articular surface for the radius. On the<br />

proximal surface, the radius facet is triangular to pyriform in outline (Figs. 4.7A, 4.8: 5),<br />

297

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