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Boyer diss 2009 1046..

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tuberosity. This crest probably delimits the radial edge of the compartment for the digital<br />

extensor musculature. The ulnar margin of the shaft of the radus develops a distinct crest<br />

for the interosseous membrance toward its distal end. The shaft swells abruptly to<br />

accommodate the distal articular surface. The dorsoradial margin of the distal epiphysis<br />

is marked by three tuberosities that formed the bony boundaries of the three extensor<br />

tendon compartments that cross the proximal carpal joint. From radial to ulnar these<br />

tuberosities correspond to the styloid process, Lister’s tubercle, and a frequently present<br />

but unnamed tubercle (Stern, 1988). The first extensor compartment would have been<br />

located radial and ventral to the styloid process. The distal articular surface itself is<br />

triangular in distal view. The radial margin forms a vertex of the triangle, while the ulnar<br />

surface is the opposing base of the triangle. The distal articular surface is marked by a<br />

faint dorsoventral ridge reflecting the respective positions of the scaphoid and lunate on<br />

either side of it. Ulnar to the lunate area, the articular surface extends on to the radial<br />

shaft for contact with the ulna.<br />

Function.—The spherical depression of the central fossa of the radial head<br />

suggests that the radius was axially mobile, as does the corresponding morphology of the<br />

humeral capitulum. A distally positioned, dorsally elevated pronator teres muscle<br />

attachment would provide a large area and lever arm for this muscle to resist substrate<br />

reaction forces causing supination, whereas the supinator muscle, attaching along the<br />

ulnar margin of the pronator teres attachment, could resist the opposite motion. Together,<br />

these muscles, as well as biceps brachii and some of the muscles originating from the<br />

supinator crest, could stabilize or rotate the humeroradial joint. The shallow distal<br />

articular surface suggests a mobile radiocarpal joint. The orientation of the joint surface<br />

293

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