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Boyer diss 2009 1046..

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Gingerich and Gunnell (1992) published a photograph of the skeleton of P. cookei<br />

(UM 87990) being formally described in this study, mounted in a life-like posture to<br />

highlight its incorporation into the Hall of Evolution of the University of Michigan’s<br />

Exhibit Museum. It was mounted clinging to a vertical tree trunk to illustrate the<br />

interpretation of it presented by these authors in abstract form in 1987.<br />

Beard (1993a) presented results of a cladistic analysis of Archonta including<br />

various plesiadapifoms. The analysis recovered plesiadapiforms as stem dermopterans.<br />

More specifically, plesiadapids formed the outgroup to a clade he called “Eudermoptera”<br />

(=Paromomyidae + Cynocephalidae). He compared and contrasted previously described<br />

morphology of plesiadapids to that of other plesiadapiforms and archontans. One<br />

synapomorphy of the clades supported by his analyses was his interpretation of carpal<br />

configuration based on the morphology of the bones of N. intermedius (USNM 442229)<br />

(p. 137, fig. 10.8). He suggested that both N. intermedius and dermopterans had a<br />

triquetrum that contacted the scaphoid and lunate on its radial surface (see Sargis [2004]<br />

for a more thorough critique of Beard [1993a]).<br />

Runestad and Ruff (1995) tested the hypothesis of Gingerich (1976) that the<br />

“robustness” of plesiadapids limbs was evidence of terrestriality in this group, and the<br />

hypothesis of Beard (1993b) that paromomyid plesiadapiforms were gliders. They did<br />

this by regressing limb lengths against limb cross-sectional areas for comparative<br />

samples of extant gliding, nongliding arboreal, and nongliding terrestrial/fossorial<br />

rodents, and marsupials. They found that gliders had the longest limb bones relative to<br />

the cross-sectional areas of their limb bones and that nongliding terrestrialists had the<br />

relatively shortest limb bones (= the most robust limbs). They analyzed previously<br />

271

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