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Boyer diss 2009 1046..

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5321), some metapodials (MNHN R 5298, 5300, 5325, 5326, 5335-37, 5340, 5345, 5351,<br />

5358, 5368, and 5370), some proximal phalanges (MNHN R 5301, 5328, 5329, 5342,<br />

5355, 5365, and 5371), some intermediate phalanges (MNHN R 5312 and 5366), some<br />

distal phalanges (MNHN R 5310, 5312, 5317, 5377, and 5381), and the distal epiphysis<br />

of MT I (MNHN R 5306).<br />

Beard’s work documented some important morphological patterns that enhance<br />

the description and analysis of P. cookei. According to Beard, associations documented<br />

for the elements of N. intermedius, by Dr. P. Houde, who originally discovered and<br />

prepared the specimen, indicate that the pedal phalanges are longer and more robust than<br />

the manual phalanges. This was later found to be true of micromomyid (Bloch and<br />

<strong>Boyer</strong>, 2007) and paromomyid plesiadapifoms (<strong>Boyer</strong> and Bloch, 2008), although not for<br />

carpolestid plesiadapiforms (Kirk et al., 2008). N. intermedius includes the first<br />

described carpal bones for a plesiadapid, including the only described scaphoid for a<br />

plesiadapiform. Its MT I, MC I, and “pollical” proximal phalanx were also the first<br />

described. Furthermore, Beard’s comparisons of carpals between N. intermedius and P.<br />

tricuspidens revealed that they are nearly identical in structure. The above information<br />

has been used by Bloch and <strong>Boyer</strong> (2007) and <strong>Boyer</strong> and Bloch (2008) to help confirm<br />

identity of elements in accumulations of semi-articulated to merely dentally-associated<br />

plesiadapiform bones.<br />

Beard’s (1989) comparative observations strongly emphasize similarities to the<br />

extant dermopteran Cynocephalus. His structural and functional analyses demonstrated<br />

that plesiadapiforms did indeed have a capacity for axial rotation of the forearm. This<br />

information, plus the documentation of a robust, divergent pollex (e.g., p. 420, table 6; p.<br />

269

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