01.09.2014 Views

Boyer diss 2009 1046..

Boyer diss 2009 1046..

Boyer diss 2009 1046..

SHOW MORE
SHOW LESS

You also want an ePaper? Increase the reach of your titles

YUMPU automatically turns print PDFs into web optimized ePapers that Google loves.

discussion of the femur indicates that it is fundamentally different from that of fossil<br />

notharctine euprimates. However, they pointed out that some features distinguishing<br />

plesiadapids from these adapiforms unite them with lorisines. Some of the distinct<br />

features of the plesiadapid femur, mentioned by Szalay et al., include the low angle<br />

between the femoral neck and shaft, the distinct constriction of the femoral neck relative<br />

to the femoral head, a large medially projecting lesser trochanter, a distally positioned<br />

third trochanter, and an anteroposteriorly shallow patellar groove. They noted the<br />

existence of the tibia (MNHN CR 410) and fibula figured by Simons (1940), and<br />

suggested that they are too fragmentary for meaningful description: they considered<br />

Simpson’s description of the tibia of N. gidleyi as adequate. Elements of the ankle joint<br />

were mentioned again; the authors referenced Szalay and Decker (1974). Specifically,<br />

they figured an astragalus based on MNHN R 610 (p. 155, fig. 12) and a calcaneum<br />

based on MNHN R 611 (p.156, fig. 13). They listed what they considered to be four<br />

derived primate characters of the astragalus and calcaneum including a pronounced<br />

groove for the tendon of flexor digitorum fibularis on the plantar side of the calcaneal<br />

sustentaculum, an astragalus with a continuous navicular and sustentacular facet, a<br />

proximodistally long tibial trochlea and a “helical-shape” to the posterior<br />

astragalocalcaneal articulation. They also provided tables (p. 158, table I; p. 160, table II)<br />

that compare these tarsal bones more systematically and thoroughly among what they<br />

consider to be condylarth, palaeoryctoid, primate, and adapid “morphotypes.” They<br />

concluded with a functional assessment, suggesting that the forearm was habitually<br />

flexed and axially mobile and that the “upper hindlimb” had a capacity for powerful<br />

extension, but a lack of leaping adaptations. They reiterated functional inferences from<br />

261

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!