Boyer diss 2009 1046..

Boyer diss 2009 1046.. Boyer diss 2009 1046..

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specimen with skull at the MNHN, but could not locate the referred specimen and was not able to see the counter slab to the type, which is housed at a separate museum. Napier and Walker (1967) commented on the locomotion of plesiadapids based on what had so far been illustrated. They suggested that plesiadapids were treeshrew or tree squirrel-like in their locomotion. Szalay (1972: p. 34) calculated brachial (93), crural (93), and intermembral (81) indices from figures provided in Simons (1964) of the Cernay material. However, it should be noted that the full length of the tibia is not actually known for P. tricuspidens, and thus the crural and intermembral indices are inferences. My own calculations, using the femur (MNHN R 408) and fragmentary tibial shaft (MNHN R 410), yield a crural index of about 92-93. Szalay and Decker (1974: figs. 3-5, 8-12) described and illustrated an astragalus (AMNH 89533) and calcaneum (AMNH 89534) of Plesiadapis cf. P. gidleyi from the early late Paleocene Saddle Locality. The Saddle locality was later determined to preserve P. anceps and N. gazini (Gingerich, 1976) and it therefore seems that these tarsal specimens should now be referred to one of these two taxa. The authors also illustrated an astragalus (no number provided) and a calcaneum MNHN R 611 of P. tricuspidens. I was able to identify the astragalus figured in this paper when I visited the MNHN. It was not, however, associated with a number. Szalay and Decker’s figure 6 reveals what appears to be the number “47” written on the specimen. These numbers have since been worn away. Russell (1964, p. 291) listed “MNHN R 5347” as an astragalus of P. tricuspidens. This fact appears to solve the mystery of the specimen’s identification. Szalay and Decker (1974) indicated that the articulation between the 258

astragalus and calcaneum was highly mobile and would permit a substantial degree of inversion and eversion, as required in an arboreal setting where substrates may occur at random orientations and descent of large tree trunks is often necessary, requiring hind foot reversal. The mid-tarsal joint was also identified as a point of axial mobility and mobility in plantar- and dorsiflexion. Szalay et al. (1975) were the first to provide substantial descriptions, figures, and analysis for P. tricuspidens material from the Cernay collection listed by Russell (1964). They also figured (p. 140, fig. 1) additional vertebral specimens of N. gidleyi (AMNH 17379), beyond those illustrated by Simpson (1935); however, these illustrations are not faithful to the actual preserved morphology and instead represent reconstructions of hypothetical complete, undistorted elements. They considered the anatomy of the vertebrae to be lacking the appropriate comparative context and thus uninformative for phylogenetic or functional considerations. They figured several humeri including MNHN BR-3-L (p. 141, fig. 2; p. 143, fig. 4) and a reconstruction based on MNHN BR- 3-L, and BR-4-L (p.144, fig. 5). Szalay et al. also considered morphology of N. gidleyi and P. walbeckensis in their discussions of humeral morphology. They concluded that the spherical, rather than cylindrical capitulum rendered Plesiadapis and euprimates unusually similar. Otherwise, they noted that the plesiadapid humeri were more similar to those of arctocyonids than to those of Paleocene “insectivorans.” They figured ulnae on pages 141-142 (figs. 2, 3). No numbers were given but I have determined that the four ulnae in these figures, from left to right in their figure, correspond to MNHN BR-7-L, MNHN R 452, MNHN R 1521, and MNHN R 443. Szalay et al. also illustrated a reconstruction of a complete (minus the styloid process) and undistorted ulna based on 259

astragalus and calcaneum was highly mobile and would permit a substantial degree of<br />

inversion and eversion, as required in an arboreal setting where substrates may occur at<br />

random orientations and descent of large tree trunks is often necessary, requiring hind<br />

foot reversal. The mid-tarsal joint was also identified as a point of axial mobility and<br />

mobility in plantar- and dorsiflexion.<br />

Szalay et al. (1975) were the first to provide substantial descriptions, figures, and<br />

analysis for P. tricuspidens material from the Cernay collection listed by Russell (1964).<br />

They also figured (p. 140, fig. 1) additional vertebral specimens of N. gidleyi (AMNH<br />

17379), beyond those illustrated by Simpson (1935); however, these illustrations are not<br />

faithful to the actual preserved morphology and instead represent reconstructions of<br />

hypothetical complete, undistorted elements. They considered the anatomy of the<br />

vertebrae to be lacking the appropriate comparative context and thus uninformative for<br />

phylogenetic or functional considerations. They figured several humeri including<br />

MNHN BR-3-L (p. 141, fig. 2; p. 143, fig. 4) and a reconstruction based on MNHN BR-<br />

3-L, and BR-4-L (p.144, fig. 5). Szalay et al. also considered morphology of N. gidleyi<br />

and P. walbeckensis in their discussions of humeral morphology. They concluded that<br />

the spherical, rather than cylindrical capitulum rendered Plesiadapis and euprimates<br />

unusually similar. Otherwise, they noted that the plesiadapid humeri were more similar<br />

to those of arctocyonids than to those of Paleocene “insectivorans.” They figured ulnae<br />

on pages 141-142 (figs. 2, 3). No numbers were given but I have determined that the four<br />

ulnae in these figures, from left to right in their figure, correspond to MNHN BR-7-L,<br />

MNHN R 452, MNHN R 1521, and MNHN R 443. Szalay et al. also illustrated a<br />

reconstruction of a complete (minus the styloid process) and undistorted ulna based on<br />

259

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