Boyer diss 2009 1046..
Boyer diss 2009 1046.. Boyer diss 2009 1046..
specimen with skull at the MNHN, but could not locate the referred specimen and was not able to see the counter slab to the type, which is housed at a separate museum. Napier and Walker (1967) commented on the locomotion of plesiadapids based on what had so far been illustrated. They suggested that plesiadapids were treeshrew or tree squirrel-like in their locomotion. Szalay (1972: p. 34) calculated brachial (93), crural (93), and intermembral (81) indices from figures provided in Simons (1964) of the Cernay material. However, it should be noted that the full length of the tibia is not actually known for P. tricuspidens, and thus the crural and intermembral indices are inferences. My own calculations, using the femur (MNHN R 408) and fragmentary tibial shaft (MNHN R 410), yield a crural index of about 92-93. Szalay and Decker (1974: figs. 3-5, 8-12) described and illustrated an astragalus (AMNH 89533) and calcaneum (AMNH 89534) of Plesiadapis cf. P. gidleyi from the early late Paleocene Saddle Locality. The Saddle locality was later determined to preserve P. anceps and N. gazini (Gingerich, 1976) and it therefore seems that these tarsal specimens should now be referred to one of these two taxa. The authors also illustrated an astragalus (no number provided) and a calcaneum MNHN R 611 of P. tricuspidens. I was able to identify the astragalus figured in this paper when I visited the MNHN. It was not, however, associated with a number. Szalay and Decker’s figure 6 reveals what appears to be the number “47” written on the specimen. These numbers have since been worn away. Russell (1964, p. 291) listed “MNHN R 5347” as an astragalus of P. tricuspidens. This fact appears to solve the mystery of the specimen’s identification. Szalay and Decker (1974) indicated that the articulation between the 258
astragalus and calcaneum was highly mobile and would permit a substantial degree of inversion and eversion, as required in an arboreal setting where substrates may occur at random orientations and descent of large tree trunks is often necessary, requiring hind foot reversal. The mid-tarsal joint was also identified as a point of axial mobility and mobility in plantar- and dorsiflexion. Szalay et al. (1975) were the first to provide substantial descriptions, figures, and analysis for P. tricuspidens material from the Cernay collection listed by Russell (1964). They also figured (p. 140, fig. 1) additional vertebral specimens of N. gidleyi (AMNH 17379), beyond those illustrated by Simpson (1935); however, these illustrations are not faithful to the actual preserved morphology and instead represent reconstructions of hypothetical complete, undistorted elements. They considered the anatomy of the vertebrae to be lacking the appropriate comparative context and thus uninformative for phylogenetic or functional considerations. They figured several humeri including MNHN BR-3-L (p. 141, fig. 2; p. 143, fig. 4) and a reconstruction based on MNHN BR- 3-L, and BR-4-L (p.144, fig. 5). Szalay et al. also considered morphology of N. gidleyi and P. walbeckensis in their discussions of humeral morphology. They concluded that the spherical, rather than cylindrical capitulum rendered Plesiadapis and euprimates unusually similar. Otherwise, they noted that the plesiadapid humeri were more similar to those of arctocyonids than to those of Paleocene “insectivorans.” They figured ulnae on pages 141-142 (figs. 2, 3). No numbers were given but I have determined that the four ulnae in these figures, from left to right in their figure, correspond to MNHN BR-7-L, MNHN R 452, MNHN R 1521, and MNHN R 443. Szalay et al. also illustrated a reconstruction of a complete (minus the styloid process) and undistorted ulna based on 259
- Page 235 and 236: Lower premolar molarization As indi
- Page 237 and 238: SUMMARY AND CONCLUSION The skull of
- Page 239 and 240: REFERENCES Bloch, J.I., Boyer, D.M.
- Page 241 and 242: TABLES Table 3.1. List of anatomica
- Page 243 and 244: Table 3.2. Anatomical abbreviations
- Page 245 and 246: Table 3.3. Size comparison among pl
- Page 247 and 248: Table 3.4 continued. European plesi
- Page 249 and 250: Figure 3.1. Cranium of Plesiadapis
- Page 251 and 252: Figure 3.3. Right maxillary teeth (
- Page 253 and 254: Figure 3.4. Cranium of Plesiadapis
- Page 255 and 256: Figure 3.5. Cranium of Plesiadapis
- Page 257 and 258: Figure 3.6. Cranium of Plesiadapis
- Page 259 and 260: Figure 3.8. Fragment from right nuc
- Page 261 and 262: Figure 3.9. Right promontorium of P
- Page 263 and 264: Figure 3.10. Cranium of Plesiadapis
- Page 265 and 266: Figure 3.12. Right dentary of Plesi
- Page 267 and 268: Figure 3.14. A, Plot of relief inde
- Page 269 and 270: CHAPTER 4: THE FIRST KNOWN SKELETON
- Page 271 and 272: among plesiadapiforms (e.g., Szalay
- Page 273 and 274: Institutional and collections abbre
- Page 275 and 276: CaL - capitulum (of humerus) antero
- Page 277 and 278: HSV - head shape variable = ln(DEW/
- Page 279 and 280: MSD - mid-shaft dorsoventral or ant
- Page 281 and 282: Ry - ray (as in “digit ray”) S-
- Page 283 and 284: History of descriptive study of the
- Page 285: illustrations of this material, exc
- Page 289 and 290: discussion of the femur indicates t
- Page 291 and 292: supinator crests. He also noted tha
- Page 293 and 294: that it may not even be an archonta
- Page 295 and 296: unstudied material. Specifically, h
- Page 297 and 298: 5321), some metapodials (MNHN R 529
- Page 299 and 300: Gingerich and Gunnell (1992) publis
- Page 301 and 302: prehensility they provide, is an in
- Page 303 and 304: euarchontans (Fig. 1.1). Their anal
- Page 305 and 306: for comparison. These include isola
- Page 307 and 308: plesiadapid samples have the same m
- Page 309 and 310: Organization of results Each bone i
- Page 311 and 312: Bloch and Boyer (2002) and N. inter
- Page 313 and 314: clavicle reflects some basic aspect
- Page 315 and 316: Humerus Description.—The right an
- Page 317 and 318: epicondyle actually projects somewh
- Page 319 and 320: cookei is absolutely longer than an
- Page 321 and 322: tuberosity. This crest probably del
- Page 323 and 324: olecranon process to estimate its t
- Page 325 and 326: distinct, convex distal radial face
- Page 327 and 328: of the midcarpal joint), and its pr
- Page 329 and 330: (there is no evidence for more than
- Page 331 and 332: matches the opposing facet on the t
- Page 333 and 334: mobility at the trapezoid-trapezium
- Page 335 and 336: Function.—The three proximal carp
astragalus and calcaneum was highly mobile and would permit a substantial degree of<br />
inversion and eversion, as required in an arboreal setting where substrates may occur at<br />
random orientations and descent of large tree trunks is often necessary, requiring hind<br />
foot reversal. The mid-tarsal joint was also identified as a point of axial mobility and<br />
mobility in plantar- and dorsiflexion.<br />
Szalay et al. (1975) were the first to provide substantial descriptions, figures, and<br />
analysis for P. tricuspidens material from the Cernay collection listed by Russell (1964).<br />
They also figured (p. 140, fig. 1) additional vertebral specimens of N. gidleyi (AMNH<br />
17379), beyond those illustrated by Simpson (1935); however, these illustrations are not<br />
faithful to the actual preserved morphology and instead represent reconstructions of<br />
hypothetical complete, undistorted elements. They considered the anatomy of the<br />
vertebrae to be lacking the appropriate comparative context and thus uninformative for<br />
phylogenetic or functional considerations. They figured several humeri including<br />
MNHN BR-3-L (p. 141, fig. 2; p. 143, fig. 4) and a reconstruction based on MNHN BR-<br />
3-L, and BR-4-L (p.144, fig. 5). Szalay et al. also considered morphology of N. gidleyi<br />
and P. walbeckensis in their discussions of humeral morphology. They concluded that<br />
the spherical, rather than cylindrical capitulum rendered Plesiadapis and euprimates<br />
unusually similar. Otherwise, they noted that the plesiadapid humeri were more similar<br />
to those of arctocyonids than to those of Paleocene “insectivorans.” They figured ulnae<br />
on pages 141-142 (figs. 2, 3). No numbers were given but I have determined that the four<br />
ulnae in these figures, from left to right in their figure, correspond to MNHN BR-7-L,<br />
MNHN R 452, MNHN R 1521, and MNHN R 443. Szalay et al. also illustrated a<br />
reconstruction of a complete (minus the styloid process) and undistorted ulna based on<br />
259